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BiologyPseudoceanic (Ref. 4066). Occurs in neritic fish community over the continental shelf (Ref. 42649). Nyctoepipelagic at the surface (Ref. 4479). Eggs of this species have been reported immediately beneath the water surface (Ref. 9212). Lipid content is 20 % in fresh body weight (Ref. 8966). Processed into fishmeal (Ref. 4066).