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"Umbrella: to 18 cm in diameter; mesoglea thin; exumbrella smooth. Marlia: 16, alternating with 1-3 marginal tentacles. Tentacles: 16-48, laterally compressed, with abaxial nematocyst warts along length. Oral arms: 4, frilled, curtainlike, length about equal to bell diameter, lip margin studded with nematocyst papillae. Gonads: 4, everted, saclike. Gastrovascular canals: total 32-96; rhopaliar canals: 16, with interjacent 1-5 tentacular canals between each rhopalium; all canals: proximally unbranched for innter 1/2-2/3 of length, distally anastomosing in an irregular network. Ring canal: near umbrellar margin. Color: according to color photographs by Rybakov , umbrella: colorless; tentacles: whitish; and gastrodermis of stomach and of oral arms: reddish-orange."