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 Thallus: thin, warted-areolate, rimose or rarely endosubstratal; areoles: scattered or contiguous, strongly convex or with irregular surface, 0.05-0.25 mm in diam., often confluent or forming a rimose crust with areoles 0.3-0.6 mm in diam.; surface: grayish beige to pale yellow; dull, esorediate; cortex: poorly developed, 5-10 µm high; medulla: mostly lacking; algal layer: 30-60 µm high; Apothecia: rounded to somewhat deformed, often in groups of 3-5, sessile, with a weakly constricted base or appressed, 0.15-0.45 mm wide; disc: ochre to reddish brown, sometimes with a pink hue due to a fine, white pruina, plane or weakly convex; margin: weakly prominent and soon excluded or lacking from the beginning,; exciple: ochre to reddish brown at the outer edge or near the hymenium, hyaline or pale yellow within, laterally 15-30 µm wide, basally 15-50(-65) µm wide, composed of branched and anastomosing, radiating hyphae with 0.7-1 µm wide lumina and apical cells with lumina 3-6 µm wide; epihymenium: yellowish to reddish brown, composed of paraphysal tips and ±sparse granules, 5-10 µm high; hymenium: hyaline, 5060(-70) µm high; paraphyses: basally hyaline, with lumina 0.7-1.2 µm wide, apically mostly with pale brown walls and lumina 2-4.5 µm wide, weakly to moderately branched (mainly below apical cells) and anastomosing; subhymenium: hyaline, 15-55 µm thick; hypothecium: hyaline or pale yellow, 0-100 µm thick; asci: clavate, Lecidella-type with an I+ blue tholus containing a lighter blue, broadly rounded axial body, 8-spored; ascospores: hyaline, simple, broadly ellipsoid, (7-)8.6-11.1(-13.5) x (4-)4.9-6.1(-7) µm; Pycnidia: not seen; Spot tests: thallus K-, C± orange, P-, UV± dark orange; Secondary metabolites: atranorin and xanthones.; Substrate and ecology: on bark, often on twigs of deciduous trees and shrubs, rarely on wood, between 50 and 500 m (one collection from 2550 m); World distribution: North America and Europe; Sonoran distribution: Baja California, southern California, and Durango.; Note: The minute, pale apothecia of Lecidea varians distinguish it from all other species treated here. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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