Description

 Thallus: crustose, coarsely granular-papillate, often strongly verrucose, coarsely cracked-areolate, rimose or areolate, thin or rather thick; prothallus: absent; areoles: 0.3-0.6 mm wide; surface: whitish gray, yellowish green, or yellowish white, dull or weakly glossy; Apothecia: on thallus, 0.5-1.5 mm in diam., sessile, slightly constricted at base; disc: black, plane or later convex, not pruinose or rarely pruinose; margin: lecideine, distinct, thin (about 0.075 mm), becoming excluded; exciple: black (at edge, reddish within) or greenish black, without algal cells; parathecium: reddish brown or yellowish brown (above, darker below), without crystals; epihymenium: green, blackish green, without or rarely with crystals; hymenium: hyaline, 55-95 µm tall, not inspersed; paraphyses: simple, rarely anastomosing or branched, slightly thickened apically; hypothecium: reddish brown; asci: clavate, lecanoral, 8-spored; ascospores: hyaline, simple, broadly ellipsoid, ellipsoid, or ovoid, (7-)9-18 x 5-9 µm; Pycnidia: not seen; Spot tests: thallus surface: K+ pale yellow or K-, C+ yellowish red (orange), KC+ yellowish red (orange), P+ yellowish or P-; Secondary metabolites: 2,5,7-trichloro-3-O-methylnorlichexanthone, 5,7-dichloro-3-O-methylnorlichexanthone, atranorin, isoarthothelin (= 2,5,7-trichloronorlichexanthone), or thiophanic acid (= 2,4,5,7-tetrachloronorlichexanthone).; Substrate and ecology: on bryophytes; World distribution: temperate Asia, Europe, and North America; Sonoran distribution: Arizona.