Characteristics

Putative autapomorphies for Boloria include (Simonsen 2005, 2006):

• Uncus bifid
• Dorsal wall of bursa ostium/antrum raised vertically with a sclerotized, arched plate just posterior to the insertion of ductus seminalis
• Bursa ostium/antrum with sclerotized ventral plates
  

Putative synapomorphies for the subgenus Boloria and include (Simonsen 2005):

• Setae on uncus oriented towards the base of uncus
  
• Uncus with pronounced ventral ridges
• Distal margin of male valve clearly concave

Putative autapomorphies for the subgenus Boloria include (Simonsen 2005, 2006):

• Phallus with two posterior, dorso-lateral areas with strongly sclerotized microtrichia
• Each half of the male juxta in dorsal view outwards pointed spatulate with convex inner margin

Clossiana include (Simonsen 2005, 2006):

• A pronounced dent present dorsally where uncus and tegumen join
• A large, membranous centro-diagonal area present on the inner surface of the male valve
• Female abdominal venter 8 with two large, weakly sclerotized, scale covered latero-posterior areas