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Putative autapomorphies for Boloria include (Simonsen 2005, 2006):

  • Uncus bifid
  • Dorsal wall of bursa ostium/antrum raised vertically with a sclerotized, arched plate just posterior to the insertion of ductus seminalis
  • Bursa ostium/antrum with sclerotized ventral plates

Putative synapomorphies for the subgenus Boloria and include (Simonsen 2005):

  • Setae on uncus oriented towards the base of uncus
  • Uncus with pronounced ventral ridges
  • Distal margin of male valve clearly concave

Putative autapomorphies for the subgenus Boloria include (Simonsen 2005, 2006):

  • Phallus with two posterior, dorso-lateral areas with strongly sclerotized microtrichia
  • Each half of the male juxta in dorsal view outwards pointed spatulate with convex inner margin

Clossiana include (Simonsen 2005, 2006):

  • A pronounced dent present dorsally where uncus and tegumen join
  • A large, membranous centro-diagonal area present on the inner surface of the male valve
  • Female abdominal venter 8 with two large, weakly sclerotized, scale covered latero-posterior areas


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Source: LepTree

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