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Vomerine teeth present. Posterior part of the tongue free and forked. Toes webbed. Omosternum and sternum ossified. Pupil of the eye horizontal. Snout moderately sharp. Male vocal sacs reduced. Shins (knee to ankle) shorter than body by 1.75-2.4 times; when the shins are positioned perpendicularly to the body axis, the heels contact or slightly overlap. When the hind leg is stretched along the body, the tibio-tarsal articulation does not usually reach the eye. Inner metatarsal tubercle small, 2.3-5.6 times shorter than first toe. Dorsal coloration greyish or grey-brown with small dark spots. Temporal spot large. Light middorsal band with distinct edges extends from eye to cloaca. Flank and thigh skin granular; granulae often red. Belly white or white-yellowish with large, irregular, partially fused blood-red spots. The red spots may alternate with dark spots, and the red pattern on the belly starts to form in about the second year of life. Males differ from females by having dark nuptial pads on the first finger.


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