Adult morphology Head as wide as or slightly wider than body, wider than long; snout subacuminate in dorsal view, rounded in lateral profile (snout deep, top of snout essentially parallel with border of lip when viewed from side); nostrils weakly protuberant, directed laterally (slightly dorsolaterally in large females); canthus rostralis distinct, its edge rounded, straight; loreal region very slightly concave, sloping abruptly to lips; lips weakly flared in large females; upper eyelid width 73.3-100.0 % of interorbital distance, bearing a few non-pungent tubercles on posterior one-third; no cranial crests; tympanum distinct, superficial, only the uppermost edge of annulus concealed by supratympanic fold; supratympanic fold prominent, ending in a subconical tubercle dorsal to insertion of forearm; tympanum virtually round, its length 46.7-64.1 % of eye length, separated from eye by a distance of less than one-half its diameter; postrictal tubercles large, subconical; skin on rest of dorsal surfaces of head finely shagreened to smooth with low tubercles scattered on occiput; choanae small, not concealed by palatal shelf of maxillary arch, slightly wider than long; vomerine odontophores low, oval in outline, situated between choanae, anterior edge of odontophore posterior to or at a line drawn through centers of choanae; each odontophore about size of a choana, bearing a row of 3-6 teeth, separated medially by a distance equal to an odontophore width; in smaller specimens, odontophores smaller, more widely separated, and appearing to be slightly more posteriad to choanae; tongue longer than wide (oval), its posterior border not notched, posterior 1/4 not adherent to floor of mouth; long vocal slits posterolateral to tongue in adult males. Skin of dorsum shagreened and bearing few other tubercles (1-2 pair above scapulae, some smaller ones on flanks); indefinite skin fold from top of supratympanic fold to about level of sacrum (lower than a dorsolateral fold); skin of upper surfaces of limbs like dorsum except shagreening finer; skin of lower surfaces smooth; discoidal folds prominent, well anteriad to groin; skin below vent areolate; vent not enclosed into sheath; no ulnar tubercles; palmar tubercle divided into minute outer and large median tubercle, median twice size of oval thenar tubercle; supernumerary palmar tubercle pungent, subconical; subarticular tubercles conical, large; fingers long and slender, bearing lateral keels (keel most notable on median surface of II); tips of digits swollen forming truncate discs on III and IV; tips of I and II rounded apically, scarcely swollen; circumferential grooves evident distally on III and IV, no evidence of ventral pads on I or II; when adpressed equally, I longer than II; thumbs of adult males bearing white, glandular nuptial pads. No tubercles or folds on heel or tarsus; inner metatarsal tubercle twice as long as wide, not compressed, approximately 1% to 2 times size of round, subconcial outer; 2-4 prominent pungent, non-conical supernumerary plantar tubercles, in addition to those at bases of toes II-IV, one (largest) in middle of sole about equidistant between base of toe IV and outer metatarsal tubercle; subarticular tubercles large, conical; toes lacking webbing, with very narrow lateral keels; toes long and slender, tips slightly dilated forming discs (smaller and less truncate than those of outer fingers); heels overlapping when flexed hind legs held at right angles to sagittal plane; hind legs short, when hind limb adpressed to body, heel reaching to a point between anterior edge of eye and midpoint between eye and nostril; shank 44.8-56.8 % of the snout-vent length. Brown above with ill-defined darker markings (interorbital bar, hourglass mark between occiput and sacrum, suprainguinal blotches); canthal - supratympanic stripe dark brown, very distinct; side of head darker than top of head, labial bars ill-defined; dark postaxillary patch, otherwise flanks not marked; limb bars about as wide as interspaces, slightly oblique; throat, breast, sides of venter, and undersides of limbs densely stippled with brown; center of belly not pigmented; posterior surfaces of thighs dark brown. In life, is tan-brown to brown with darker brown to black markings. The ventral surfaces are gray to grayish brown. The concealed surfaces of the hind limbs are light brown with a slight orange wash to the groin and behind the knee. The undersides of the limbs are flesh-colored. The iris is chocolate brown.