The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • C. Michael Hogan & World Wildlife Fund. 2015. Highveld grasslands. Encyclopedia of Earth. National Council for Science and Environment. Washington DC
  • Channing, A. (2001). Amphibians of Central and Southern Africa. Comstock books in herpetology. x, 470 p., [24] p. of plates. Ithaca, N.Y.: Comstock Pub. Associates.
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • J.P.H. Acocks. 1988. Veld types of South Africa. Memoirs of the Botanical Survey of South Africa 57: 1-146. (An update of the first edition published in 1953),
  • Orrell T. (custodian) (2013). ITIS Global: The Integrated Taxonomic Information System (version Sept 12). In: Species 2000 & ITIS Catalogue of Life, 11th March 2013 (Roskov Y., Kunze T., Paglinawan L., Orrell T., Nicolson D., Culham A., Bailly N., Kirk P., Bourgoin T., Baillargeon G., Hernandez F., De Wever A., eds). Digital resource at Species 2000: Reading, UK.
  • [Anonymous] (2004). Atlas and Red Data Book of the Frogs of South Africa, Lesotho and Swaziland. (MinterL., BurgerM., HarrisonJ., BraackH H., BishopP J., KloepferD., Ed.).SI/MAB Series. 9, Washington, D.C.: Smithsonian Institution.


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