128465Trachypithecus francoisiEndangered (EN)<i>Trachypithecus francoisi</i> (Pousargues, 1898)Francois' langurFrancois's leaf monkeyIndochinese LangurdeTonkin-SchwarzlangurenFranÁois' leaf monkeyenFranÁois's langurenFranÁois's leaf monkeyenFrancois' langurenFrançois' leaf monkeyenFrancois'LangurenFrançois's langurenFrançois's leaf monkeyenIndochinese LangurenTonkin leaf monkeyesLangur de francoisfrLangur de franÁoisfrLangur de françoisfrSemnopithèque de franÁoisfrSemnopithèque de françoisfrTrachypithecus francoisiitTrachypithecus francoisinlFrançois' langoerruЛангур или трахипитек ФрансуаTate, G. 1947. Mammals of Eastern Asia. New York: The Macmillan Company.15587115587327198false1Helin, S., N. Ohtaishi, L. Houji. 1999. The Mammalian of China. Beijing: China Forestry Publishing House.17676117676327198false1Glover, A. 1938. The Mammals of China and Mongolia. New york: American Museum of Natural History.17677117677327198false1Becher, F., J. Nijboer, J. van der Kuilen, A. Beynen. 2001. Chemical analysis and consistency of faeces produced by captive monkeys (Francois langurs, Trachypithecus francoisi) fed supplemental fibre. Veterinary Quarterly, 23/2: 76-80.17678117678327198false1Ding, B., L. Hai-peng, Z. Ya-ping, L. Zimin, Y. Wei. 2000. Taxonomic Status of the White-head Langur (Trachypithecus francoisi leucoscephalus) Inferred from Allozyme Electrophoresis and Random Amplified Polymorphism DNA (RAPD). Zoological Studies, 39/4: 313-318.17679117679327198false1Nowak, R. 1997. "Walker's Mammals of the World Online 5.1" (On-line). Accessed October 12, 2004 at http://www.press.jhu.edu/books/walkers_mammals_of_the_world/primates/primates.cercopithecidae.trachypithecus.html .17680117680327198false1Collins, L., M. Roberts. 1978. Arboreal folivores in captivity- maintenance of a delicate minority. Pp. 5-12 in G.G. Montgomery, ed. The Ecology of Arboreal Folivores. Washington: Smithsonian Institute.17681117681327198false1Davies, A. 1994. Colobine populations. Pp. 285-310 in J.F. Oates, ed. Colobine Monkeys: Their Ecology Behavior and Evolution. Cambridge: Cambridge University Press.17682117682327198false1Isbell, L. 1994. Predation on primates: ecological patterns and evolutionary consequences. Evolutionary Anthropology, 3: 151-154.17683117683327198false1Yeager, C., K. Kool. 2000. Behavioral ecology of Asian colobines. Pp. 496-521 in P. Whitehead, J. Clifford, eds. Old World Monkeys. Cambridge: Cambridge University Press.17684117684327198false1Massicot, P. 2004. "Animal Info- Francois' Leaf Monkey" (On-line). Accessed October 12, 2004 at http://www.animalinfo.org/species/primate/tracfran.htm .17685117685327198false1Hussain, A., V. Shanmugam, V. Bhullar, B. Beer, D. Vallet. 2003. Screening for simian foamy virus infection by using a combined antigen Western blot assay: evidence for a wide distribution among Old World primates and identification of four new divergent viruses. Virology, 306/2: 248-257.17686117686327198false1Tate, G. 1947. Mammals of Eastern Asia. New York: The Macmillan Company.155871155871077290false1Helin, S., N. Ohtaishi, L. Houji. 1999. The Mammalian of China. Beijing: China Forestry Publishing House.176761176761077290false1Glover, A. 1938. The Mammals of China and Mongolia. New york: American Museum of Natural History.176771176771077290false1Becher, F., J. Nijboer, J. van der Kuilen, A. Beynen. 2001. Chemical analysis and consistency of faeces produced by captive monkeys (Francois langurs, Trachypithecus francoisi) fed supplemental fibre. Veterinary Quarterly, 23/2: 76-80.176781176781077290false1Ding, B., L. Hai-peng, Z. Ya-ping, L. Zimin, Y. Wei. 2000. Taxonomic Status of the White-head Langur (Trachypithecus francoisi leucoscephalus) Inferred from Allozyme Electrophoresis and Random Amplified Polymorphism DNA (RAPD). Zoological Studies, 39/4: 313-318.176791176791077290false1Nowak, R. 1997. "Walker's Mammals of the World Online 5.1" (On-line). Accessed October 12, 2004 at http://www.press.jhu.edu/books/walkers_mammals_of_the_world/primates/primates.cercopithecidae.trachypithecus.html .176801176801077290false1Collins, L., M. Roberts. 1978. Arboreal folivores in captivity- maintenance of a delicate minority. Pp. 5-12 in G.G. Montgomery, ed. The Ecology of Arboreal Folivores. Washington: Smithsonian Institute.176811176811077290false1Davies, A. 1994. Colobine populations. Pp. 285-310 in J.F. Oates, ed. Colobine Monkeys: Their Ecology Behavior and Evolution. Cambridge: Cambridge University Press.176821176821077290false1Isbell, L. 1994. Predation on primates: ecological patterns and evolutionary consequences. Evolutionary Anthropology, 3: 151-154.176831176831077290false1Yeager, C., K. Kool. 2000. Behavioral ecology of Asian colobines. Pp. 496-521 in P. Whitehead, J. Clifford, eds. Old World Monkeys. Cambridge: Cambridge University Press.176841176841077290false1Massicot, P. 2004. "Animal Info- Francois' Leaf Monkey" (On-line). Accessed October 12, 2004 at http://www.animalinfo.org/species/primate/tracfran.htm .176851176851077290false1Hussain, A., V. Shanmugam, V. Bhullar, B. Beer, D. Vallet. 2003. Screening for simian foamy virus infection by using a combined antigen Western blot assay: evidence for a wide distribution among Old World primates and identification of four new divergent viruses. Virology, 306/2: 248-257.176861176861077290false1Tate, G. 1947. Mammals of Eastern Asia. New York: The Macmillan Company.155871155871141128false1Helin, S., N. Ohtaishi, L. Houji. 1999. The Mammalian of China. Beijing: China Forestry Publishing House.176761176761141128false1Glover, A. 1938. The Mammals of China and Mongolia. New york: American Museum of Natural History.176771176771141128false1Becher, F., J. Nijboer, J. van der Kuilen, A. Beynen. 2001. Chemical analysis and consistency of faeces produced by captive monkeys (Francois langurs, Trachypithecus francoisi) fed supplemental fibre. Veterinary Quarterly, 23/2: 76-80.176781176781141128false1Ding, B., L. Hai-peng, Z. Ya-ping, L. Zimin, Y. Wei. 2000. Taxonomic Status of the White-head Langur (Trachypithecus francoisi leucoscephalus) Inferred from Allozyme Electrophoresis and Random Amplified Polymorphism DNA (RAPD). Zoological Studies, 39/4: 313-318.176791176791141128false1Nowak, R. 1997. "Walker's Mammals of the World Online 5.1" (On-line). Accessed October 12, 2004 at http://www.press.jhu.edu/books/walkers_mammals_of_the_world/primates/primates.cercopithecidae.trachypithecus.html .176801176801141128false1Collins, L., M. Roberts. 1978. Arboreal folivores in captivity- maintenance of a delicate minority. Pp. 5-12 in G.G. Montgomery, ed. The Ecology of Arboreal Folivores. Washington: Smithsonian Institute.176811176811141128false1Davies, A. 1994. Colobine populations. Pp. 285-310 in J.F. Oates, ed. Colobine Monkeys: Their Ecology Behavior and Evolution. Cambridge: Cambridge University Press.176821176821141128false1Isbell, L. 1994. Predation on primates: ecological patterns and evolutionary consequences. Evolutionary Anthropology, 3: 151-154.176831176831141128false1Yeager, C., K. Kool. 2000. Behavioral ecology of Asian colobines. Pp. 496-521 in P. Whitehead, J. Clifford, eds. Old World Monkeys. Cambridge: Cambridge University Press.176841176841141128false1Massicot, P. 2004. "Animal Info- Francois' Leaf Monkey" (On-line). Accessed October 12, 2004 at http://www.animalinfo.org/species/primate/tracfran.htm .176851176851141128false1Hussain, A., V. Shanmugam, V. Bhullar, B. Beer, D. Vallet. 2003. Screening for simian foamy virus infection by using a combined antigen Western blot assay: evidence for a wide distribution among Old World primates and identification of four new divergent viruses. Virology, 306/2: 248-257.176861176861141128false11. IUCN Red List (April, 2006) <a href="http://www.iucnredlist.org">http://www.iucnredlist.org</a>68888716888871420485false12. Richardson., M. (2006) <i>Pers. Comm.</i>69293116929311420485false13. Human Ageing Genomic Resources – AnAge Database (April, 2006) <a href="http://genomics.senescence.info/species/index.html">http://genomics.senescence.info/species/index.html</a>69293216929321420485false14. CITES (April, 2006) <a href="http://www.cites.org">http://www.cites.org</a>69293316929331420485false15. The Primata (April, 2006) <a href="http://www.theprimata.com/trachypithecus_francoisi.html">http://www.theprimata.com/trachypithecus_francoisi.html</a>69293416929341420485false16. Honolulu Zoo (April, 2006) <a href="http://www.honoluluzoo.org/francois%5Fmonkey.htm">http://www.honoluluzoo.org/francois%5Fmonkey.htm</a>69293516929351420485false17. Animal Info (April, 2006) <a href="http://www.animalinfo.org/species/primate/tracfran.htm">http://www.animalinfo.org/species/primate/tracfran.htm</a>69293616929361420485false18. UNEP-WCMC (April, 2006) <a href="http://www.unep-wcmc.org">http://www.unep-wcmc.org</a>69293716929371420485false19. Institute for Ecology and Biological Resources – Primate Conservation in Vietnam (April, 2006) <a href="http://coombs.anu.edu.au/~vern/iebr.html">http://coombs.anu.edu.au/~vern/iebr.html</a>69293816929381420485false1Richard Weigl (2005) Longevity of Mammals in Captivity; from the Living Collections of the World. Kleine Senckenberg-Reihe 48: Stuttgart.73092417309242391827false1The highly folivorous diet of Francois' langur led to the evolution of enlarged salivary glands and a sacculated stomach, which combine to encourage break down and digestion of the tough fibre in leaves, with the help of bacteria living in the upper chamber of the stomach. The lower chamber contains acid as in most mammals, whereas the upper chamber is neutral to create more favourable conditions for the bacteria. Francois' langur is not restricted to eating leaves; it also consumes fruit, seeds, nectar, shoots and insects, varying its diet with seasonal changes in the abundance of these foods (5). Fleshy immature leaves are preferred (5), preventing the langurs from needing to drink frequently (6). The group moves as it feeds, ending each day in a new place. They settle down to sleep at dusk, preferring large trees in good weather, or limestone caves if it's cold or raining (8).</p><p>An active and noisy species, members of troops indulge in daily mutual grooming in rest periods between feeding (5) (6). The group consists of 4 – 27 (usually around a dozen) individuals and led by the females, who operate a reasonably changeable hierarchy amongst themselves, particularly when it comes to caring for the young (2) (5) (6). There is usually just one adult male in the group who will not participate in caring for the young, except to respond to their distress calls. He will also rarely groom other members of the group, but expects to be groomed himself. Females share responsibility for the infants, who are born singly once a year. They are immediately cared for by 'allomothers' as well as their own mother, who will nurse the young monkey for up to two years before weaning. Once weaned, the bond between mother and infant is no different than between any other members of the group. After three or four years juveniles become sexually mature, and will commonly leave to join another group or to form an all-male bachelor group. Francois' langurs are not fully grown until the age of six or seven (5) (6).943cdc0e4c581ed1e4b3cedcd97d035c2609311WildscreenCopyright Wildscreen 2003-2008ARKiveCopyrightCopyright Wildscreen 2003-2008. Some rights reservedhttp://creativecommons.org/licenses/by-nc-sa/3.0/SupplierARKivehttp://www.arkive.org/5978arkiveLocationSource URLView original data objecthttp://www.arkive.org/francois-langur/trachypithecus-francoisi/CitationEnglishcc-by-nc-sa 3.0Texthttp://content10.eol.org/content_large.jpghttp://content1.eol.org/content_medium.jpghttp://content2.eol.org/content_small.jpg13290302267232704204128529783328662512953009717226298227131AnimaliaNOT EVALUATEDAnimalia694ChordataNOT EVALUATEDChordata1642MammaliaNOT EVALUATEDMammalia1645PrimatesNOT EVALUATEDPrimates1657CercopithecidaeNOT EVALUATEDCercopithecidae15096TrachypithecusNOT EVALUATEDTrachypithecus128465Trachypithecus francoisiEndangered (EN)<i>Trachypithecus francoisi</i> (Pousargues, 1898)Portland State University, MA Education
7klansBA Physics - Johns Hopkins University
MS Education - Johns Hopkins University
ME Engineering Physics - University of Virginia
10peterPh.D., University of Bristol (1976)
D.Sc., Queen's University, Belfast (1990)
11paddydeveloper13pleary11793wilsondI am currently a PhD candidate in the Department of Environmental Science, Policy, and Management at the University of California-Berkeley.
List of publications:
Rothman, J.M., C.A. Chapman, D. Twinomugisha, M.D. Wasserman, J.E. Lambert, and T.L. Goldberg. 2008. Measuring physical traits of primates remotely: the use of parallel lasers. American Journal of Primatology 70: 1191-1195.
Snaith,T.V., C.A. Chapman, J.M. Rothman, and M.D. Wasserman. 2008. Bigger groups have fewer parasites and similar cortisol levels: a multi-group analysis in red colobus monkeys. American Journal of Primatology 70: 1072-1080.
Chapman, C.A., L. Naughton-Treves, M.J. Lawes, M.D. Wasserman, and T.R. Gillespie. 2007. Population declines of colobus in western Uganda and conservation value of forest fragments. International Journal of Primatology 28: 513-528.
Chapman, C.A., M.D. Wasserman, T.R. Gillespie, M.L. Speirs, M.J. Lawes, T.L. Saj, and T.E. Ziegler. 2006. Do nutrition, parasitism, and stress have synergistic effects on red colobus populations living in forest fragments? American Journal of Physical Anthropology 131: 525-534.
Chapman, C.A., M.D. Wasserman, and T.R. Gillespie. 2006. Behavioural patterns of colobus in logged and unlogged forests: The conservation value of harvested forests. In: E. Newton-Fisher, H. Notman, V. Reynolds, J.D. Patterson (eds.). Primates of Western Uganda. Pp. 373-390. Springer, New York.
Chapman, C.A., T. Webb, R. Fronstin, M.D. Wasserman, and A.M. Santamaria. 2005. Assessing dietary protein of colobus monkeys through fecal sample analysis: A tool to evaluate habitat quality. African Journal of Ecology 43: 276-278.
Wasserman, M.D., and Chapman, C.A. 2003. Determinants of colobine monkey abundance: The importance of food energy, protein and fiber content. Journal of Animal Ecology 72: 650-659.
Member of:
International Primatological Society – 2005 to present
Society for Conservation Biology – 2005 to present
American Society of Primatologists – 2007 to present
Association for Tropical Biology & Conservation – 2009 to present.20368mwassermAsst. Project Manager, Antarctic Invertebrates, Smithsonian Institution
Visiting Scientist, Chemicals Affecting Insect Behavior Lab, US Agricultural Research Service, 2005-2006
Hammock, J., Vinyard, B., Dickens, J. 2007. Response to host plant odors and aggregation pheromone by larvae of the Colorado potato beetle on a servosphere. Arthropod-Plant Interactions, 1(1):27-35
PhD, Biological Oceanography, Massachussetts Institute of Technology/Woods Hole Oceangraphic Institution, 200520470jhammockUniversity of Michigan, PhD 1997
Thesis: Social behavior and vocal communication of American Crows (Corvus brachyrhynchos)
Member of Ecological Society of America, Entomological Society of America.25567csparr33422mstuderCurator of Zoology (Fishes), Field Museum of Natural History34360mwestneatEOL Species Pages Coordinator, Ph. D. in Entomology, University of Arizona 1999, Diploma in Biology, Freie Universität Berlin 198935200KatjaWilliam Miller, PhD
Assistant Professor
Baker University
Dept. of Biology
PO Box 65
Baldwin city, Kansas
66006-0065
35243WMillerUniversity of California Berkeley
35508tuco36197MarthaTestuserB.A. Zoology, M.S. Embryology/physiology
college faculty for 44 years
member HAPS (Human Anatomy and Physiology Society)36558judyjunkyardActive Member of Wildlife society of Africa, herpetological society of southern Africa and general manager and curator of reptile research and information center (Zebra Country lodge, Cullinan district Pretoria)37046VaughanScientist National Institutes of Health38259saimirifaculty, Department of Biology
University of Miami
Coral Gables, Fl 33124
AAAS, ICRS, Sigma Xi
http://www.bio.miami.edu/Fac/Sealey.html38707ksealeyAssociate Professor of Biology, Curator of Mammals and Birds, University of Wisconsin - Stevens Point
Yahnke CJ, 2006. Habitat use and natural history of small mammals in the Paraguayan chaco. Mastozoologia Neotropical, 13(1);103-116.
Yahnke CJ, 2006. Testing optimal foraging theory using bird predation on goldenrod galls. American Biology Teacher, 68(8):471-475.
Yahnke CJ, PL Meserve, TG Ksiazek, and JN Mills. 2001. Patterns of infection with Laguna Negra virus in wild populations of Calomys laucha in the central Paraguayan chaco. American Journal of Tropical Medicine and Hygiene, 65(6):768-776.
Yahnke CJ, I Gamarra de Fox, and F Colman. 1998. Mammalian species richness in Paraguay: the effectiveness of national parks in preserving biodiversity. Biological Conservation 84:263-268.
Yahnke CJ, J Unger, BA Lohr, DA Meritt, and W Heuschele. 1997. Age specific fecundity, litter size, and sex ratio in the chacoan peccary (Catagonus wagneri). Zoo Biology 16:301-307.
Yahnke, CJ, WE Johnson, E Geffen, D Smith, F Hertel, MS Roy, CF Bonacic, TK Fuller, B Van Valkenburgh, and RK Wayne. 1996. Darwin’s fox: a distinct endangered species in a vanishing habitat. Conservation Biology 10:366-375.
Yahnke CJ. 1995. Metachromism and the insight of Wilfred Osgood: evidence of common ancestry for Darwin’s fox and the Sechura fox. Revista Chilena de Historia Natural 68:459-467
American Society of Mammalogists
National Association of Biology Teachers
Sigma XI
39446cyahnkeEmeritus Fellow, Trinity College Dublin
Holdich, Catherine, David M., Noel, Pierre Y., Reynolds, Julian D. and Haffner, Patrick (eds) (2006). Atlas of crayfish in Europe. Museum national d'Histoire naturelle, Paris, 187 pages. Patrimoines naturels, 64.
Hudson, Anne V. and Reynolds, Julian D. (1984). Distribution of Irish intertidal Talitridae. Bulletin of the Irish biogeographical Society, 8, 63-76.
Reynolds, Julian D. (1976). Occurrence of the fresh-water Bryozoan, Cristatella mucedo Cuvier, in British Columbia. Syesis, 9, 365-366.
Smyth, Thomas and Reynolds, Julian D. (1995). Survival ability of statoblasts of freshwater Bryozoa found in Renvyle Lough, County Galway. Biology and Environment: Proceedings of the Royal Irish Academy 95B (1), 65-68.
Wickenberg, Maria and Reynolds, Julian D. (2002). A recent Irish record of the woodlouse Acaeroplastes melanurus (Budde-Lund, 1885) (Isopoda: Porcellionidae), considered to be extinct in the British Isles. Bulletin of the Irish Biogeographical Society, 26, 60-63.39544jrynoldsAffiliated with EOL Species Pages Group39552lshapiroStaff in a natural history museum
Member of a professional society (SECEM)39803eliadelomCurator of marine invertebrates, National Museums Northern Ireland (Ulster Museum)
Author Sponges of the British Isles - A colour guide and working document, 1992 Edition. Ackers, R.G., Moss, D. & Picton, B. E. 1992. Marine Conservation Society, UK. Revised and extended, 2007, Bernard Picton, Christine Morrow & Rob van Soest. PDf and website40160BernardPicton030