The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Lehtonen S. 2011. Towards Resolving the Complete Fern Tree of Life. PLoS ONE 6(10): e24851. doi:10.1371/journal.pone.0024851
  • Pryer, K.M., E. Schuettpelz, P.G. Wolf, et al. 2004. Phylogeny and evolution of ferns (monilophytes) with a focus on the early leptosporangiate divergences. American Journal of Botany 91(10): 1582-1598.
  • Pryer, K.M., H. Schneider H, A.R. Smith, et al. 2001. Horsetails and ferns are a monophyletic group and the closest living relatives to seed plants. Nature 409: 618–622.
  • Smith, A.R., K.M. Pryer, E. Schuettpelz, et al. 2006. A classification of extant ferns. Taxon 55(3): 705-731.


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