The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Larsen, T. H. (2004). Dung beetles / escarabajos peloteros. (VriesendorpC., Rivera ChavezL., MoskovitsD., ShoplandJ., Ed.).Peru: Megantoni. Rapid Biological Inventories Report. Report 15, 77-84, 185-192. Chicago, Illinois: The Field Museum.
  • Larsen, T. H., & Forsyth A. (Submitted). Dung Beetles of the Upper Amazon. [Book about the ecology, natural history, distribution, identification and conservation of Scarabaeine dung beetles of the southwest Amazon in Peru and Bolivia, based on unpublished research by the authors and many other researchers.].
  • Redtenbacher L. 1867. Reise der Osterreichischen Fregatte Novara um die Erde in den Jahren 1857-58-59 unter der Befehlen des Commodore B.von Wullerstorf-Urbair. Zoologie II part. Coleopteren. Wien (1868) :1-249
  • Schoolmeesters P. (2015). Scarabs: World Scarabaeidae Database (version Jul 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.


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