The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.
- Harrison J.Du G. & Scholtz C.H. & Chown S.L. 2003. A revision of the endemic south-western African dung beetle subgenus Scarabaeus (Pachysoma), including notes on other flightless Scarabaeini. Journal of Natural History 37:305-355
- Schoolmeesters P. (2013). Scarabs: World Scarabaeidae Database (version Aug 2012). In: Species 2000 & ITIS Catalogue of Life, 11th March 2013 (Roskov Y., Kunze T., Paglinawan L., Orrell T., Nicolson D., Culham A., Bailly N., Kirk P., Bourgoin T., Baillargeon G., Hernandez F., De Wever A., eds). Digital resource at www.catalogueoflife.org/col/. Species 2000: Reading, UK.
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