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Overview
Comprehensive Description
Description
General: Grass Family (Poaceae). Koleagrass is an introduced, tall, robust, rapid-developing bunchgrass with short rhizomes. This species includes both Phalaris tuberosa var. stenoptera (previously known as Harding grass) and Phalaris tuberosa var. hirtiglumis (previously known as koleagrass). P.t. var. hirtiglumis was recognized to have bigger seeds, more vigorous seedlings, yellow-green leaves, and hairy glumes. Both of these varieties have been submerged taxonomically into Phalaris aquatica.
Distribution: Koleagrass was introduced from Morocco. Used for pasture and erosion control in California and sparingly in other parts of the southwestern U.S. Consult the PLANTS Web site for its U.S. distribution.
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Alternative names
Bulbous canarygrass, hardinggrass, Phalaris tuberosa var. hirtiglumis (koleagrass), Phalaris tuberosa var. stenoptera (Harding grass), Towoomba canary grass, alpiste bulbeaux, phalaris tubereux, herbe de Harding, knolliges Glanzgras, capim-doce, rabillode cordero
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Distribution
Algeria (Africa & Madagascar)
Argentina (South America)
Australia (Oceania)
Baleares (Europe)
Bolivia (South America)
Brazil (South America)
Chile (South America)
New Zealand (Oceania)
Ecuador (South America)
Malta (Europe)
Morocco (Africa & Madagascar)
Cyprus (Asia)
Ethiopia (Africa & Madagascar)
France (Europe)
Greece (Europe)
Iraq (Asia)
Israel (Asia)
Italy (Europe)
Kenya (Africa & Madagascar)
Libya (Africa & Madagascar)
Madeira (Africa & Madagascar)
Egypt (Africa & Madagascar)
Mexico (Mesoamerica)
South Africa (Africa & Madagascar)
United States (North America)
Canada (North America)
Uruguay (South America)
Peru (South America)
Venezuela (South America)
Corsica (Europe)
Yugoslavia (Europe)
Portugal (Europe)
Canary Islands (Africa & Madagascar)
Syria (Asia)
Spain (Europe)
Turkey (Asia)
United Kingdom (Europe)
Pakistan (Asia)
China (Asia)
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Forzza, R. C. & et al. 2010. 2010 Lista de espécies Flora do Brasil. http://floradobrasil.jbrj.gov.br/2010/.
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Marticorena, C. & M. Quezada. 1985. Catálogo de la Flora Vascular de Chile. Gayana, Bot. 42: 1–157.
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Soreng, R. J., G. Davidse, P. M. Peterson, F. O. Zuloaga, E. J. Judziewicz, T. S. Filgueiras & O. Morrone. 2003 and onwards. On-line taxonomic novelties and updates, distributional additions and corrections, and editorial changes since the four published volumes of the Catalogue of New World Grasses (Poaceae) published in Contr. U.S. Natl. Herb. vols. 39, 41, 46, and 48. http://www.tropicos.org/Project/CNWG:. In R. J. Soreng, G. Davidse, P. M. Peterson, F. O. Zuloaga, T. S. Filgueiras, E. J. Judziewicz & O. Morrone Internet Cat. New World Grasses. Missouri Botanical Garden, St. Louis.
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Herrera Arrieta, Y. & A. Cortés Ortiz. 2010. Listado florístico y aspectos ecológicos de la familia Poaceae para Chihuahua, Durango y Zacatecas, México. J. Bot. Res. Inst. Texas 4(2): 711–738.
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Tovar, Ó. 1993. Las Gramíneas (Poaceae) del Perú. Ruizia 13: 1–480.
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Espejo Serna, A., A. R. López-Ferrari & J. Valdés-Reyna. 2000. Poaceae. Monocot. Mexic. Sinopsis Floríst. 10: 7–236 [and index].
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Edgar, E., M. A. O'Brien & H. E. Connor. 1991. Checklist of pooid grasses naturalised in New Zealand. 1. Tribes Nardeae, Stipeae, Hainardieae, Meliceae, and Aveneae. New Zealand J. Bot. 29: 101–116.
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Gibbs Russell, G. E., W. G. Welman, E. Reitief, K. L. Immelman, G. Germishuizen, B. J. Pienaar, M. v. Wyk & A. Nicholas. 1987. List of species of southern African plants. Mem. Bot. Surv. S. Africa 2(1–2): 1–152(pt. 1), 1–270(pt. 2).
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Hokche, O., P. E. Berry & O. Huber. 2008. Nuev. Cat. Fl. Vas. Venezuela 1–860. Fundación Instituto Botánico de Venezuela, Caracas, Venezuela.
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Jørgensen, P. M. & S. León-Yánez. (eds.) 1999. Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75: i–viii, 1–1181.
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Luteyn, J. L. 1999. Páramos, a checklist of plant diversity, geographical distribution, and botanical literature. Mem. New York Bot. Gard. 84: viii–xv, 1–278.
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Munz, P. A. 1974. Fl. S. Calif. 1–1086. University of California Press, Berkeley.
http://www.tropicos.org/Reference/1719
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Munz, P. A. 1968. Suppl. Calif. Fl. 1–224. University of California Press, Berkeley.
http://www.tropicos.org/Reference/1718
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Flora of China Editorial Committee. 2006. Fl. China 22: 1–733. Science Press & Missouri Botanical Garden Press, Beijing & St. Louis.
http://www.tropicos.org/Reference/1029690
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Zuloaga, F. O., E. G. Nicora, Z. E. R. Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella. 1994. Catálogo de la familia Poaceae en la República Argentina. Monogr. Syst. Bot. Missouri Bot. Gard. 47: i–xi, 1–178.
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Renvoize, S. A. 1998. Gram. Bolivia i–xxx, 1–644. The Royal Botanic Gardens, Kew.
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Sharp, D. & B. K. Simon. 2002. AusGrass: Grasses of Australia. CD-ROM, Version 1.0. CD–ROM.
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Covas, G. 1970. Hordeum, Phalaris. In: A. L. Cabrera (ed.), Gramíneas. 4(2): 175–189. In A. L. Cabrera Fl. Prov. Buenos Aires. Instituto Nacional de Tecnología Agropecuaria, Buenos Aires.
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Baldini, R. M. 1995. Revision of the genus Phalaris L. (Gramineae). Webbia 49(2): 265–329.
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Soreng, R. J. 2003. Phalaris. In Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae. Contr. U.S. Natl. Herb. 48: 479–488.
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Anderson, D. E. 1961. Taxonomy and distribution of the genus Phalaris. Iowa State Coll. J. Sci. 36(1): 1–96.
http://www.tropicos.org/Reference/9811
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Algeria (Africa & Madagascar)
France (Europe)
Italy (Europe)
Morocco (Africa & Madagascar)
Poland (Europe)
Tunisia (Africa & Madagascar)
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Baldini, R. M. 1995. Revision of the genus Phalaris L. (Gramineae). Webbia 49(2): 265–329.
http://www.tropicos.org/Reference/1004089
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Argentina (South America)
Brazil (South America)
Uruguay (South America)
United States (North America)
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Zuloaga, F. O., E. G. Nicora, Z. E. R. Agrasar, O. Morrone, J. Pensiero & A. M. Cialdella. 1994. Catálogo de la familia Poaceae en la República Argentina. Monogr. Syst. Bot. Missouri Bot. Gard. 47: i–xi, 1–178.
http://www.tropicos.org/Reference/43109
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Smith, L. B., D. C. Wasshausen & R. M. Klein. 1982. Gramíneas. Gêneros: 45. Deschampsia até 84. Pseudechinolaena. 1(GRAM): 443–906. In R. Reitz Fl. Il. Catarin. Herbário "Barbarosa Rodrigues", Itajaí, Brasil.
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Burkart, A. 1969. Gramíneas. 2: 1–551. In A. Burkart Fl. Il. Entre Ríos. Instituto Nacional de Tecnología Agropecuaria, Buenos Aires.
http://www.tropicos.org/Reference/19863
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SPECIMEN BASED RECORD. Published protolog data.
http://www.tropicos.org/Reference/9990002
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Australia (Oceania)
Mexico (Mesoamerica)
Uruguay (South America)
United States (North America)
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Beetle, A. A. 1977. Noteworthy grasses from Mexico V. Phytologia 37(4): 317–407.
http://www.tropicos.org/Reference/2538
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Munz, P. A. & D. D. Keck. 1959. Cal. Fl. 1–1681. University of California Press, Berkeley.
http://www.tropicos.org/Reference/1717
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Rosengurtt, B., B. R. A. Maffei & P. I. Artucio. 1970. Gram. Urug. [i–vii], 1–489. Universidad de la República, Montevideo.
http://www.tropicos.org/Reference/19689
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National Distribution
United States
Origin: Exotic
Regularity: Regularly occurring
Currently: Unknown/Undetermined
Confidence: Confident
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Global Range: Native to Mediterrean Europe. Introduced to CA.
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Distribution
- Flora of Pakistan @ eFloras.org
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Adaptation
Koleagrass is adapted in California below 2,000 feet to the Mediterranean-like climatic zone or where frost heaving is not a problem. It will grow where there is 12 inches of annual rainfall and there is soil profile development (clay layer). Moisture holding capacity of the soil becomes less critical as rainfall increases. It establishes itself early, grows well during cold winter months, and reseeds readily.
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Physical Description
Morphology
Comments
Bulb Canary Grass is introduced in Pakistan and is a valuable fodder grass.
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Physical Description
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Description
- Flora of Pakistan @ eFloras.org
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Comments
- Flora of China @ eFloras.org
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Description
- Flora of China @ eFloras.org
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Ecology
Habitat
Habitat & Distribution
- Flora of China @ eFloras.org
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Dispersal
Establishment
Seedbed Preparation: Good stands require a spring-summer fallow to reduce competition from annuals in fall. Fall seeding is done on a firm, weed-free seedbed prepared after the first effective rains (1.5-2.0 inches). This ensures adequate moisture to support seedling growth and destroys the first crop of volunteer annuals. If fallow is prepared before weed species produce seed, the fall weed competition is reduced, but not eliminated. Fall seeding can follow a summer clean-up crop of Sudangrass (Sorghum bicolor ssp. drummondii). Planting is accomplished directly in the Sudangrass stubble.
Method of Seeding: Best results are obtained by drilling the seed. Use of a common grain drill is possible if the seed is first mixed with rice hulls as a dilutor and the drill is properly calibrated. If seed is broadcast and covered by harrowing or dragging, the seeding rate is increased 1.5 times. Seeding depth should be ¼-1/2 inch in all cases.
Weed Control: During the establishment season, mowing may be necessary to reduce competition from annuals. Set the mower at a height compatible with regrowth of the koleagrass and other perennials. Remove as little leaf area of the koleagrass and other perennials as possible so that development will not be retarded.
Fertilization: Do not apply fertilizer in the establishment year. In the second and subsequent years, 200 pounds per acre 16-20-0 (ammonium-phosphate-sulfate) is recommended in the fall.
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Life History and Behavior
Cyclicity
Flower/Fruit
- Flora of Pakistan @ eFloras.org
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Molecular Biology and Genetics
Barcode
Locations of barcode samples
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Statistics of barcoding coverage
| Specimen Records: | 5 |
| Specimens with Sequences: | 0 |
| Specimens with Barcodes: | 0 |
| Public Records: | 0 |
| Species: | 1 |
| Species With Barcodes: | 0 |
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Conservation
Conservation Status
National NatureServe Conservation Status
United States
Rounded National Status Rank: NNA - Not Applicable
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NatureServe Conservation Status
Rounded Global Status Rank: GNR - Not Yet Ranked
Reasons: Native to Mediterrean Europe and introduced to CA.
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Status
Please consult the PLANTS Web site and your State Department of Natural Resources for this plant’s current status (e.g. threatened or endangered species, state noxious status, and wetland indicator values).
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Threats
Management
Cultivars, improved and selected materials (and area of origin)
‘Perla’ Cultivar- This cultivar was introduced from Morocco. It is easy to establish, grows well during cold winter months, and produces good seed crops. Stands frequently improve through natural reseeding. Also, it’s adapted to soils with restricting layer in Mediterranean climatic zone wherever average annual rainfall is 400mm or more.
Contact your local Natural Resources Conservation Service (formerly Soil Conservation Service) office for more information. Look in the phone book under ”United States Government.” The Natural Resources Conservation Service will be listed under the subheading “Department of Agriculture.”
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Seed production
Koleagrass can be grown as an annual for purposes of seed production. At the Lockeford Plant Materials Center, irrigated rows spaced 30 inches apart yield 400-600 pounds per acre of seed when harvested with a binder in late spring following fall seeding. Koleagrass seed averages 267,000 seeds per pound.
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Control
Please contact your local agricultural extension specialist or county weed specialist to learn what works best in your area and how to use it safely. Always read label and safety instructions for each control method. Trade names and control measures appear in this document only to provide specific information. USDA, NRCS does not guarantee or warranty the products and control methods named, and other products may be equally effective.
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Management
Koleagrass should not be grazed until completion of growth the first season. At the end of the green feed period, dry forage can be grazed if the plants are large enough to resist pulling by livestock. An average stubble height of three to four inches should be maintained.
Established Stands: Koleagrass starts growth in fall coincident with the first rains. Mature stands of koleagrass should be ready for grazing (6-8 inches high) by early January. The amount of growth depends on the amount of rain, its distribution, temperature and soil fertility. Cold tolerance is improved by the inclusion of phosphorus with nitrogen during fall fertilization. Grass can be grazed at range readiness without harm to the plant so long as the ground is firm and not so wet that damage from trampling might occur. Grazing should stop when fertilized annual range is ready for use or when there is a three-inch average stubble height. Grazing will normally end in late February. However, lack of moisture or poor growing conditions may require earlier termination. Infrequently, when abundant rainfall occurs in late winter, grazing can continue through mid-March. Following winter grazing, the perennials, including koleagrass, must be rested to allow recovery and re-growth. This deferment period should continue until the annual range grasses begin to dry up and lose their high feed value. By the time the annual range is dry, koleagrass will have produced seed and stored food reserves in the roots. The leaves will still be green and will remain green for about one month after the annuals are dry. This palatable forage can be grazed heavily until the average stubble height is three inches. When grazing ends, many plants will be heavily used, others partially grazed, and there will be numerous seed stalks left.
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Weediness
This plant may become weedy or invasive in some regions or habitats and may displace desirable vegetation if not properly managed. Please consult with your local NRCS Field Office, Cooperative Extension Service office, or state natural resource or agriculture department regarding its status and use. Weed information is also available from the PLANTS Web site. Please consult the Related Web Sites on the Plant Profile for this species for further information.
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Relevance to Humans and Ecosystems
Benefits
Uses
Koleagrass was developed for improvement of annual grass range. Its forage production is typically no greater than can be expected from native range. However, this perennial produces forage earlier in the season and stays green longer. Fertilized Koleagrass can extend the green feed period two months or more in the winter compared to unfertilized native grasses, and can add about one month to the green feed period in spring when moisture is not limiting. NOTE: This species has been known to have toxic effects on sheep and cattle.
Koleagrass is now considered the standard perennial grass for improvement of annual grass range in California. Please consult the Related Web Sites on the Plant Profile for further information on this species.
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Wikipedia
Phalaris aquatica
Phalaris aquatica, syn. P. tuberosa, known by the common names Bulbous canarygrass and Harding grass, is a species of grass in the genus Phalaris of the Poaceae Family.
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Description
It is an erect, waist-high, stout perennial bunch grass with grayish to bluish green leaves. Flowering heads are dense, spike-like, and usually two to five inches long. It is slow to develop from seed, but can form large bunches after several years.[2]
Phalaris arundinacea ('Reed canary grass') differs from Harding grass in having more distinct rhizomes and an inflorescence that is compact at first but later becomes more open as the branches spread.
Hybrids of Harding grass and reed canary grass have been produced. Varieties include 'AQ1', 'Uneta', and 'Australis'.
Toxicity
Some Phalaris species contain gramine, which can cause brain damage, other organ damage, central nervous system damage and death in sheep.[3]
Leaves and seedlings contain the tryptamine hallucinogens DMT, 5-MeO-DMT and related compounds.[4] A raw, dried plant Phalaris aquatica contains approximately 0.1% DMT, 0.022% 5-MeO-DMT, and 0.005% bufotenin.[5] A particular strain of P. aquatica from Italy, labeled 'AQ-1', was reported to contain in excess of 1.0% alkaloid concentration.[6]
Invasive species
Harding Grass is an invasive species in grassland, oak woodland, and chaparral, and riparian habitats. The California native grasses in rangelands and native grasslands are affected.
See also
References
- ^ "Phalaris aquatica information from NPGS/GRIN". www.ars-grin.gov. http://www.ars-grin.gov/cgi-bin/npgs/html/taxon.pl?27511. Retrieved 2008-04-02.
- ^ ucce.ucdavis.edu
- ^ Toxicants of Plant Origin - Google Book Search. books.google.com. http://books.google.com/books?hl=en&lr=&ie=UTF-8&id=eASgQyXq8xMC&oi=fnd&pg=PA169&dq=%22mimosa+hostilis%22+dimethyltryptamine&ots=pe6oz-oODM&sig=D6kaAQYmEqSOkJ4kkztPcm2K10I#PPA171,M1. Retrieved 2008-04-20.
- ^ Tryptamine Carriers FAQ
- ^ Phalaris / DMT FAQ
- ^ Ayahuasca: alkaloids, plants & analogs: assembled by Keeper of the Trout
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