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Overview
Brief Summary
The Lebanon Cedar (Cedrus libani) resembles a number of other Cedrus cedars. Cedrus deodara has pendulous leading shoot and branch tips (i.e., the entire new shoot gently droops), whereas C. libani has upright, stiff leading shoots, occasionally with just the branch tips drooping. Cedrus atlantica has densely pubescent (downy) blue leaves, whereas C. libani has glabrous (smooth) or sparsely pubescent green leaves. (Choukas-Bradley and Alexander 1987; Cope 2001)
The natural habitat of Cedrus libanii in the mountains of Lebanon has been substantially impacted by humans over centuries. Intensive logging for ship building and construction, as well as land-clearing for agriculture, were recorded as early as the 3rd millennium B.C. Vanishing forests were already mentioned during the 1st century B.C. and depletion of these forests has continued. It is now estimated that the current 2,000 hectares of patchy cedar forests found in Lebanon are the remnants of more than 500,000 hectares of post-glacial forest. In Turkey, cedar forests cover almost 110,000 hectares and occur primarily in the Taurus mountains, the steep slopes of which have somewhat sheltered its forests from overexploitation and extirpation. (Fady et al. 2008 and references therein)
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Distribution
Distribution
Lebanon Cedar is native to the eastern Mediterranean region, including Turkey, Lebanon, and Syria (Choukas-Bradley 1987; Bird 1994; Kurt et al. 2008). It is more widely planted as a highly regarded ornamental (Dirr 1998).
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Physical Description
Morphology
Morphology
The Lebanon Cedar has a thick, massive trunk and very wide-spreading branches, the lower ones sweeping the ground. The dark green, densely packed leaves are borne in horizontal tiers. Young trees are slender pyramids, but become flat-topped as they mature. (Bird 1994; Dirr 1998)
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Size
Size
Mature height is about 40 meters (a typical 10-year-old specimen would be around 6 meters tall) (Bird 1994).
After 40 to 70 years, Lebanon Cedar reaches 12 to 18 meters, but it can grow to a maximum size of about 23 to 37 meters in height with a 24 to 30 meter spread (Dirr 1998).
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Look Alikes
Lookalikes
Cedrus libani resembles a number of other Cedrus cedars. Cedrus deodara has pendulous leading shoot and branch tips (i.e., the entire new shoot gently droops), whereas C. libani has upright, stiff leading shoots, occasionally with just the branch tips drooping. Cedrus atlantica has densely pubescent (downy) blue leaves, whereas C. libani has glabrous (smooth) or sparsely pubescent green leaves. (Choukas-Bradley and Alexander 1987; Cope 2001)
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Ecology
Habitat
Habitat
Lebanon Cedar is currently found in the mountains of Turkey, Syria, and Lebanon from 1,400 to 2,200 meters above sea level (Quezel and Medail 2003, cited in Fady et al. 2008).
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Associations
Associations
fruitbody of Ganoderma lucidum is saprobic on dead stump of Cedrus libani
Other: minor host/prey
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Life History and Behavior
Life Expectancy
Life Expectancy
The largest specimens of Lebanon Cedar are several hundred years old, but the tree grows surprisingly quickly in its early years--around 15 cm each year for around 70 years--then slows down (Bird 1994).
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Evolution and Systematics
Evolution
Phylogeography
Fady et al. (2008) studied the genetic variation of Cedrus libani across Turkey and Lebanon. Substantial genetic variation was detected, consistent with the general pattern that eastern Mediterranean conifers tend to harbor higher levels of genetic variation than conifer populations elsewhere (likely due to the fact that their populations during the late glacial maximum (18,000 years ago) were somewhat sheltered in favorable environments and did not suffer strong demographic and genetic bottlenecks). However, this high genetic diversity was not evenly partitioned among C. libani populations. Based on their genetic data, Fady et al. support proposals to recognize two divergent C. libani taxa, one in Lebanon and one in Turkey, which they suggest should be treated as distinct subspecies. The authors suggest that the overall lower genetic diversity found in cedars from Lebanon (as compared with Turkey) is likely the result of overexploitation in the form of logging and/or grazing.
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Systematics or Phylogenetics
Cedrus phylogeography
C. deodara, the Himalayan cedar constitutes a separate gene pool from the Mediterranean cedars. The
large genetic distances observed between C. deodara and Mediterranean cedars reflect the high geographical distance between these two regions and an ancient divergence between these taxa.
- Bou Dagher-Kharrat M., Grenier G., Bariteau M., Brown S., Siljak-Yakovlev S. et Savouré A., 2001. Karyotype analysis reveals interspecific differentiation in the Cedrus genus despite genome size and base composition constancy. Theoretical and Applied Genetics. 2001- 103: 846-854.
- Fady B., Lefèvre F., Reynaud M., Vendramin G.G., Bou-Dagher-Kharrat M., Andizei M., Pastorelli R., Savouré A., Bariteau, M., gene flow among different taxonomùic units: evidence from nuclear and cytoplasmic markers in Cedrus plantation forests. Theoretical and Applied Genetics. 2003- 107:1132-1138.
- Bou Dagher-Kharrat M., Mariette S., Fady B., Lefevre F., Grenier G., Plomion C. et Savouré A., Geographical diversity and genetic relationships among Cedrus species assayed by AFLP. 2007. Tree Genetics & Genomes 3: 275-285
Unreviewed
Cedrus phylogeography
C. deodara, the Himalayan cedar constitutes a separate gene pool from the Mediterranean cedars. The
large genetic distances observed between C. deodara and Mediterranean cedars reflect the high geographical distance between these two regions and an ancient divergence between these taxa.
- Bou Dagher-Kharrat M., Grenier G., Bariteau M., Brown S., Siljak-Yakovlev S. et Savouré A., 2001. Karyotype analysis reveals interspecific differentiation in the Cedrus genus despite genome size and base composition constancy. Theoretical and Applied Genetics. 2001- 103: 846-854.
- Fady B., Lefèvre F., Reynaud M., Vendramin G.G., Bou-Dagher-Kharrat M., Andizei M., Pastorelli R., Savouré A., Bariteau, M., gene flow among different taxonomùic units: evidence from nuclear and cytoplasmic markers in Cedrus plantation forests. Theoretical and Applied Genetics. 2003- 107:1132-1138.
- Bou Dagher-Kharrat M., Mariette S., Fady B., Lefevre F., Grenier G., Plomion C. et Savouré A., Geographical diversity and genetic relationships among Cedrus species assayed by AFLP. 2007. Tree Genetics & Genomes 3: 275-285
Unreviewed
Cedrus phylogeography
C. deodara, the Himalayan cedar constitutes a separate gene pool from the Mediterranean cedars. The
large genetic distances observed between C. deodara and Mediterranean cedars reflect the high geographical distance between these two regions and an ancient divergence between these taxa.
- Bou Dagher-Kharrat M., Grenier G., Bariteau M., Brown S., Siljak-Yakovlev S. et Savouré A., 2001. Karyotype analysis reveals interspecific differentiation in the Cedrus genus despite genome size and base composition constancy. Theoretical and Applied Genetics. 2001- 103: 846-854.
- Fady B., Lefèvre F., Reynaud M., Vendramin G.G., Bou-Dagher-Kharrat M., Andizei M., Pastorelli R., Savouré A., Bariteau, M., gene flow among different taxonomùic units: evidence from nuclear and cytoplasmic markers in Cedrus plantation forests. Theoretical and Applied Genetics. 2003- 107:1132-1138.
- Bou Dagher-Kharrat M., Mariette S., Fady B., Lefevre F., Grenier G., Plomion C. et Savouré A., Geographical diversity and genetic relationships among Cedrus species assayed by AFLP. 2007. Tree Genetics & Genomes 3: 275-285
Unreviewed
Cedrus phylogeography
C. deodara, the Himalayan cedar constitutes a separate gene pool from the Mediterranean cedars. The
large genetic distances observed between C. deodara and Mediterranean cedars reflect the high geographical distance between these two regions and an ancient divergence between these taxa.
- Bou Dagher-Kharrat M., Grenier G., Bariteau M., Brown S., Siljak-Yakovlev S. et Savouré A., 2001. Karyotype analysis reveals interspecific differentiation in the Cedrus genus despite genome size and base composition constancy. Theoretical and Applied Genetics. 2001- 103: 846-854.
- Fady B., Lefèvre F., Reynaud M., Vendramin G.G., Bou-Dagher-Kharrat M., Andizei M., Pastorelli R., Savouré A., Bariteau, M., gene flow among different taxonomùic units: evidence from nuclear and cytoplasmic markers in Cedrus plantation forests. Theoretical and Applied Genetics. 2003- 107:1132-1138.
- Bou Dagher-Kharrat M., Mariette S., Fady B., Lefevre F., Grenier G., Plomion C. et Savouré A., Geographical diversity and genetic relationships among Cedrus species assayed by AFLP. 2007. Tree Genetics & Genomes 3: 275-285
Unreviewed
Conservation
Conservation Status
IUCN Red List Assessment
Red List Category
Red List Criteria
Version
Year Assessed
- Needs updating
Assessor/s
Reviewer/s
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Conservation Status
The natural habitat of Cedrus libani in the mountains of Lebanon has been substantially impacted by humans over centuries. Intensive logging for ship building and construction, as well as land-clearing for agriculture, were recorded as early as the 3rd millennium B.C. Vanishing forests were already mentioned during the 1st century B.C. and depletion of these forests has continued. It is now estimated that the current 2,000 hectares of patchy cedar forests found in Lebanon are the remnants of more than 500,000 hectares of post-glacial forest. In Turkey, cedar forests cover almost 110,000 hectares and occur primarily in the Taurus mountains, the steep slopes of which have somewhat sheltered its forests from overexploitation and extirpation. (Fady et al. 2008 and references therein)
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Relevance to Humans and Ecosystems
Benefits
Uses
Traditional people in southern Turkey produce a wood extract, called katran, from C. libani, and use it to protect wooden structures against insects and fungi, to fight parasites and bacteria, and to heal wounds and cure various diseases in humans and domestic animals, both internally and externally. Kurt et al. (2008) discuss traditional methods of producing katran, its use by local people, and its chemical composition.
The wood of Lebanon Cedar has been greatly appreciated since ancient times for its durability, density, color, and insecticidal properties (Fady et al. 2008). According to biblical references, wood from this tree was used in the construction of King Solomon's temple (Choukas-Bradley and Alexander 1987).
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Wikipedia
Cedrus libani
Cedrus libani (Lebanon Cedar or Cedar of Lebanon), is a species of cedar native to the mountains of the Mediterranean region, in Lebanon, western Syria and south central Turkey, with varieties of it (some treated as separate species by some authors) in southwest Turkey, Cyprus, and the Atlas Mountains in Algeria and Morocco in northwest Africa.[1][2][3] The Cedars Conservancy parks in Lebanon were candidates in the New 7 Wonders of Nature competition but were eliminated at the end of the first stage while the Jeita Grotto was chosen as the Lebanese national candidate.[4]
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Description
The Lebanon Cedar is an evergreen coniferous tree growing up to 40 m (130 ft) tall, with a trunk up to 2.5 m (8 ft 2 in) diameter. The crown is conic when young, becoming broadly tabular with age with more or less level branches. The shoots are dimorphic, with long shoots and short shoots. The leaves are needle-like, spaced out on the long shoots, and in clusters of 15-45 on the short shoots; they are 5-30 mm (1⁄4–1 3⁄16 in) in length, quadrangular in cross-section, and vary from green to glaucous blue-green with stomatal bands on all four sides. The seed cones are produced often every second year, and mature in 12 months from pollination; mature cones in late autumn are 8-12 cm (3–4 3⁄4 in) long and 4-6 cm (1 1⁄2–2 3⁄8 in) wide.
Taxonomy
Cedrus libani was first described by French botanist Achille Richard. There are four ill-defined varieties, treated as subspecies or closely related species by some authors:
- Cedrus libani var. libani (Lebanon Cedar): Lebanon, western Syria and south central Turkey
- Cedrus libani var. stenocoma (Turkish Cedar): southwest Turkey
- Cedrus libani var. brevifolia (Cyprus Cedar): Cedrus libani var. atlantica (Atlas Cedar): Atlas Mountains
The treatment of the Turkish, Cyprus and Atlas Cedars as varieties or subspecies of Lebanon Cedar is found primarily in botanical and floristic works,[2][5][6][7][8][9][10][11][12] while treatment as separate species is more widespread in popular horticultural use,[13] but also in some botanical works.[1][14][15] The discrepancy in treatment derives largely from the very narrow gene base of trees in cultivation, which gives a false impression of distinctiveness of the taxa, not borne out when the wider range of variation found in wild trees is examined.[16]
Ecology
In Lebanon and Turkey it is most abundant at altitudes of 1,000-2,000 m (3,300–6,500 ft), where it forms pure forests or mixed forests with Cilician Fir (Abies cilicica), European Black Pine (Pinus nigra), and several juniper (Juniperus) species. In Cyprus, it occurs at 1,000-1,525 m (3,300–5,000 ft) (reaching the summit of Mount Paphos). In the Atlas Mountains, it occurs at 1,370–2,200 m (4,500–7,200 ft) in pure forests or mixed with Abies species and Juniperus thurifera.[1]
History, symbolism and uses
Cedar of Lebanon was important to various civilizations. The trees were used by the ancient Phoenicians for building trade and military ships, as well as houses and temples. The Egyptians used its resin for mummification, and its sawdust was found in pharaohs' tombs. The Sumerian Epic of Gilgamesh designates the cedar groves of Lebanon as the dwelling of the gods to where Gilgamesh ventured. They once burned cedar in their ceremonies. Jewish priests were ordered by Moses to use the bark of the Lebanon Cedar in circumcision and treatment of leprosy. Isaiah used the Lebanon Cedar as a metaphor for the pride of the world[17] According to the Talmud, Jews once burned Lebanese cedar wood on the Mount of Olives to announce the new year. Kings far and near requested the wood for religious and civil constructs, the most famous of which are King Solomon's Temple in Jerusalem and David's and Solomon's Palaces. It was also used by Romans, Greeks, Persians, Assyrians and Babylonians.[18]
Historically, extensive deforestation has occurred, with only small remnants of the original forests surviving. Deforestation has been particularly severe in the Lebanon and Cyprus; on Cyprus, only small trees up to 25 m (82 ft) tall survive, though Pliny the Elder recorded cedars 40 m (130 ft) tall there.[19] Extensive reforestation of cedar is carried out in the Mediterranean region, particularly Turkey, where over 50 million young cedars are being planted annually.[20] The Lebanese populations are also now expanding through a combination of replanting and protection of natural regeneration from browsing by goats.[21]
Regional significance
The Lebanon Cedar is the national emblem of Lebanon, and it is seen on the Lebanese flag. It is also the main symbol of the Cedar Revolution, along with many political parties in Lebanon such as the Kataeb, the National Liberal Party and the Lebanese Forces.[22]
As a result of long exploitation, very few old trees remain in Lebanon, but there is now an active program to conserve and regenerate the forests. The forest of the Cedars of God in Bsharri and the Barouk forest are national reserves in Lebanon.[23][24][25] Extensive replanting is taking place in Turkey, where approximately 300 square kilometres (74,000 acres) of cedar are planted annually.[21]
Horticultural use
The Lebanon Cedar is widely planted as an ornamental tree in parks and large gardens, often being planted in avenues, and as focal point trees in large lawns. The most prominent landscaping feature in London's historic Highgate Cemetery is its "Circle of Lebanon", where a Lebanon Cedar stands in the centre of a circular trench cut into the ground and lined with mausoleums.[26]
Location
Map of the cedar's forests and the surrounding mountains (www.mount-lebanon.com)
Gallery
| Wikimedia Commons has media related to: Cedrus libani |
Lebanon Cedar in Al Shouf Cedar Nature Reserve, Barouk, Lebanon | Atlas Cedar Cedrus libani var. atlantica at Kew Gardens | Cedrus libani var. atlantica 'Glauca' branches | Cedrus libani var. atlantica, Domaine de Mariemont, Belgium |
Small Lebanon Cedar in half barrel pot, Noda, Chiba Japan |
See also
References
- ^ a b c Farjon, A. (1990). Pinaceae. Drawings and Descriptions of the Genera. Koeltz Scientific Books ISBN 3-87429-298-3.
- ^ a b Greuter, W., Burdet, H. M., & Long, G. (eds.), (1984). Med-Checklist – A critical inventory of vascular plants of the circum-mediterranean countries. Cedrus libani
- ^ Conifer Specialist Group (1998). Cedrus libani. 2006. IUCN Red List of Threatened Species. IUCN 2006. www.iucnredlist.org. Retrieved on 12 May 2006.
- ^ New7wonders.com
- ^ Hooker, J. D. (1862). On the Cedars of Lebanon, Taurus, Algeria and India. Nat. Hist. Rev. 2: 11-18.
- ^ Battander, J.-A. & Trabut, L. (1905). Flora de l'Algérie.
- ^ Schwarz, O. (1944). Anatolica. Feddes Repertorium 54: 26-34.
- ^ Coode, M. J. E., & Cullen, J., eds. (1965). Pinaceae. In: Flora of Turkey and the East Aegean Islands 1: 67-85. Edinburgh University Press.
- ^ Meikle, R. D. (1977). Flora of Cyprus vol. 1. Bentham - Moxon Trust, Royal Botanic Gardens, Kew. London.
- ^ Browicz, K. & Zielinski, J. (1982). Chorology of Trees and Shrubs in southwest Asia vol. 1.
- ^ Frankis, M. & Lauria, F. (1994). The maturation and dispersal of cedar cones and seeds. International Dendrology Society Yearbook 1993: 43-46.
- ^ Güner, A., Özhatay, N., Ekim, T., & Başer, K. H. C. (ed.). 2000. Flora of Turkey and the East Aegean Islands 11 (Supplement 2): 5-6. Edinburgh University Press. ISBN 0-7486-1409-5
- ^ Walters, W. M. (1986). European Garden Flora Vol 1. ISBN 0-521- 24859-0.
- ^ Christou, K. A. (1991). The genetic and taxonomic status of Cyprus Cedar, Cedrus brevifolia (Hook.) Henry. Mediterranean Agronomic Institute of Chania, Greece.
- ^ Gymnosperm database: Cedrus.
- ^ Gaussen, H. (1964). Genre Cedrus. Les Formes Actuelles. Trav. Lab. For. Toulouse T2 V1 11: 295-320
- ^ Isaiah 2:13
- ^ History of the Cedars
- ^ Willan, R. G. N. (1990). The Cyprus Cedar. Int. Dendrol. Soc. Yearbk. 1990: 115-118.
- ^ Anon. History of Turkish Forestry. Turkish Ministry of Forestry.
- ^ a b Khuri, S., & Talhouk, S. N. (1999). Cedar of Lebanon. Pages 108-111 in Farjon, A., & Page, C. N. Status Survey and Conservation Action Plan: Conifers. IUCN/SSC Conifer Specialist Group. ISBN 2-8317-0465-0.
- ^ Lebanon, the Cedars' Land
- ^ Talhouk, S. N. & Zurayk, S. 2003. Conifer conservation in Lebanon. Acta Hort. 615: 411-414.
- ^ Semaan, M. & Haber, R. 2003. In situ conservation on Cedrus libani in Lebanon. Acta Hort. 615: 415-417.
- ^ Cedars of Lebanon Nature Reserve
- ^ Highgatecemetery.net
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