The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Ananjeva, N. & Orlov, N. 2005. Lizards of North Eurasia. Reptilia (GB) (38): 54-63.
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV. 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Ananjeva, Natalia B.;Tuniyev, Boris S. 1992. Historical biogeography of the Phrynocephalus species of the USSR. Asiatic Herpetological Research 4: 76-98.
  • BARABANOV, A.V. & N.B. ANANJEVA. 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa 1399: 1-56.
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23.
  • Bedriaga, J.V. 1909. Amphibien und Reptilien. In: Wissenschaftliche Resultate der Reisen N.M. Przewalskijs durch Zentralasien. Zoologische Teil. Band 3. Part 1. Lacertilia. (in Russian). Sankt-Petersburg, Vol 3 (1) 3: 73-102, 367.
  • Bedriaga, JACQUES VON. 1907. Neue Saurier aus Russisch-Asien. Annu. Mus. Zool. Acad. Impér. Sci. St.-Petersbourg, 10 [1905]: 210-243.
  • Bedriaga, JACQUES VON. 1907. Verzeichnis der von der Central-Asiatischen Expedition unter Stabs-Kapitän W. Roborowski in den Jahren 1893-1895 gesammelten Reptilien. Annu. Mus. Zool. Acad. Impér. Sci. St.-Petersbourg, 10 (3-4): 159-200 [1905].
  • Bedriaga. 1907. . Wiss. Res. Przewalski Central-Asien untern. Reisen 3, Zool., Abt. 1, Lief. 2: 207.
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp.
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp.
  • Eichwald, E. 1831. Zoologia specialis, quam expositis animalibus tum vivis, tum fossilibus potissimuni rossiae in universum, et poloniae in specie, in usum lectionum publicarum in Universitate Caesarea Vilnensi. Zawadski, Vilnae.
  • Elpatjevsky, VLADIMIR S. 1907. . Izvest. Taschkent Otd. russ. geogr. Obschtsch. 4 (7): 51.
  • Engelmann,W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Fitzinger, L. 1823. in Lichtenstein: in Evermann's Reise [von Orenburg] nach Buchara. E. H. G. Christiani, Berlin.
  • Gmelin, J.F. 1789. Caroli a Linné Systema naturae. 13. ed., Tom 1 Pars 3. G. E. Beer, Lipsiae. 1033-1516 pp.
  • Gmelin,S.G. 1789. In: Linnaeus, C. Systema Naturae per regna tria naturae secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus 1, Pars III. Editio decima tertia, aucta, reformata. Georg. Emanuel Beer, Lipsiae [Leipzig]. pp. 1033-1516.
  • Golubev, M.L. 1989. . Vest. Zool., 5: 38.
  • Golubev, M.L. 1991. About the name Agama ocellata Lichtenstein in Eversmann, 1823 (Reptilia, Agamidae) with redescription of the types. Herpetological Researches (1):12-17.
  • Golubev, M.L. 1993. The variegated toad agama in Djungar Gate (Eastern Kazakstan) with notes on certain systematic problems of Phrynocephalus versicolor Str. (Reptilia, Agamidae). Asiatic Herpetological Research 5: 51-58.
  • Ji, Xiang; Yue-Zhao Wang & Zheng Wang. 2009. New species of Phrynocephalus (Squamata, Agamidae) from Qinghai, Northwest China. Zootaxa 1988: 61-68.
  • Kaup, J. 1827. Zoologische Monographien. Isis von Oken 20: 610—625 [columns].
  • Lambert, M.R.K. 2002. Preliminary observations on herpetofaunal diversity in the Almaty region, Southern Kazakhstan (September 1998). Herpetological Bulletin (79): 7-13.
  • Manilo, Valantina V.;Golubev, Michael L. 1993. Karyotype information on some toad agamas of the Phrynocephalus guttatus species group (Sauria, Agamidae) of the former USSR. Asiatic Herpetological Research 5: 105-108.
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp.
  • Mezhzherin, Sergei;Golubev, Michael L. 1993. Allozyme variation and genetic relationships within the Phrynocephalus guttatus species group (Sauria: Agamidae) in the former USSR. Asiatic Herpetological Research 5: 59-64.
  • Nikolsky, A. M. 1913. Herpetologia caucasica [in Russian]. Tiflis (Museum caucasicum), 272 pp.
  • Nikolsky,A.M. 1915. Fauna Rossii sopredel. Stran, Rept. Vol. 1 [Reptiles of Russia and adjacent countries. Reptiles (Reptilia). Vol. 1. Chelonia and Sauria; in Russian]. Imper. Acad. Sci., Petrograd, 532 pp. [Translated from Russian by the Israel Program for Scientific Translations 1963].
  • Pallas,Peter S. 1827. Zoographia Rosso-Asiatica, sistens omnium animalium in extsenso imperio rossico et adjacentibus maribus observatorum recensionexu, domicilia, mores et descriptiones, anatomen atque icones plurimorum. Volumen Tertum [=volume 3]. Caes. Acad. Sci., Petropoli (= St. Petersburg). vii + 428 + cxxv pages.
  • Pang, Junfeng; Yuezhao Wang, Yang Zhong, A. Rus Hoelzel, Theodore J. Papenfuss, Xiaomao Zeng, Natalia B. Ananjeva and Ya-ping Zhang. 2003. A phylogeny of Chinese species in the genus Phrynocephalus (Agamidae) inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 27: 398-409.
  • Peters G. 1984. Die Krötenkopfagamen Zentralasiens (Agamidae: Phrynocephalus). MITTEILUNGEN AUS DEM ZOOLOGISCHEN MUSEUM IN BERLIN 60(1): 23-67.
  • Roitberg, E. S., Mazanaeva, L.F., Ilyina, E. & Orlova, V.F. 2000. Die Echsen Dagestans (Nordkaukasus, Russland): Artenliste und aktuelle Verbreitungsdaten (Reptilia: Sauria: Gekkonidae, Agamidae, Anguidae, Scincidae et Lacertidae). Faunist. Abh. Staatl. Mus. Tierk. Dresden 22 (8).
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp.
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp.
  • Tabachishin, Vasily G. and Evgeni V. Zavialov. 2005. Present distribution and taxonomic status of the Spotted Toad-headed Agama Phrynocephalus guttatus (Gmelin, 1789) in the Volga region, Russia. Herpetozoa 18 (3/4): 141-146. .
  • Terentjew & TSCHERNOW. 1949. Opredelitel Presmykatushchikhsya i Zemnovodnykh. [Encyclopedia of reptiles and amphibians, in Russian]. Moscow.
  • Wittig, W. 2002. Haltung und Nachzucht von Phrynocephalus guttatus. Elaphe 10 (3): 29-31.
  • Zeng, Xiaomao; Wang, Yaozhao; Liu, Zhijun;Yu, Pin. 1994. A COMPARATIVE STUDY ON THE KARYOTYPE OF THREE SPECIES IN Phrynocephalus (Agamidae). Acta Zootaxonomica Sinica 1994 (2).
  • Zhan, A. & Fu, J. 2009. Microsatellite DNA markers for three toad-headed lizard species (Phrynocephalus vlangalii, P. przewalskii and P. guttatus). Molecular Ecology Resources 9: 535–538.
  • Zhao,Ken-Tang. 1979. Studies on the classification and distibution of sand lizards in China (in Chinese). Acta Univ. Inner Mongolia, Hohhot 1979 (2): 111-121.


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