Sub-Saharan Africa: all S of Sahara except Namibia, W South Africa.
Zambezian Halophytics Habitat
The Makgadikgadi spiny agama (Agama hispida makarikarika) is endemic to the Makgadikgadi Pans complex within the Botswana element of the Zambezian halophytics ecoregion. This agama typically inhabits the edges of the pans but it is difficult to spot, since it buries itself in the sand during the heat of the day.
One of the largest saltpans in the world, the Makgadikgadi Pan complex in Botswana stretches out over 12,000 square kilometres. The ecoregion is classified within the Flooded Grasslands and Savanna biome. Surrounded by the semi-arid Kalahari savannas, the pans experience a harsh climate, hot with little rain, and are normally a vast, glaring expanse of salt-saturated clay. These pans are sustained by freshwater from the Nata River, and more infrequently, from input from the Okavango Alluvial Fan by way of the Boteti River. Saline- and drought-tolerant plant species generally line the pan perimeters, with grasslands further removed from the pans.
For most of the year the pans are depauperate in bird numbers, except for ostriches and species such as the Chestnut-banded sand-plover and Kittlitz’s plover (Charadrius pallidus, C. pecuarius). The sole hospitable area to birds during these times is the Nata Delta, which has a permanent water source and a small resident population of waterbirds including grebes (Podiceps spp.), cormorants (Phalacrocorax spp.), ducks and plovers (Charadrius spp.) with a few flamingos (Phoenicopterus ruber, Phoeniconaias minor) and pelicans (Pelecanus spp.). The grasslands surrounding the pans support a moderate bird fauna with species such as ostriches, secretary birds (Sagittarius serpentarius), kori bustards (Ardeotis kori), korhaans (Eupodotis spp.), sandgrouse (Pterocles spp.) and francolin (Francolinus spp.) being common. The Hyphaene palms to the west of the pans are nesting sites for, among others, the greater kestrel (Falco rupicoloides) and the palm-nut vulture (Gypohierax angolensis). After good rains the pans are transformed into a vibrant paradise, attracting thousands of waterbirds, most of which come to breed on the pans. Wattled and southern crowned cranes (Grus carunculatus, Balearica regulorum), saddle-billed, marabou and open-billed storks (Ephippiorhynchus senegalensis, Leptoptilos crumeniferus, Anastomus lamelligerus), African fish eagles (Haliaeeetus vocifer), black-necked grebes (Podiceps nigricollis), Caspian terns (Hydroprogne caspia), eastern white and pink-backed pelicans (Pelecanus onocrotalus, P. rufescens), geese and waders such as avocets (Recurvirostra avosetta), black-winged stilts (Himantopus himantopus), plovers, sandpipers and teals (Anas spp.) congregate around the pans. The most spectacular arrival are the greater and lesser flamingos (Phoenicopterus ruber and Phoeniconaias minor) that flock to the pans in their thousands.
Most mammalian taxa within the ecoregion inhabit the grasslands surrounding the pans. These include Hartebeest (Alcelaphus buselaphus), Gemsbok (Oryx gazella), Springbok (Antidorcas marsupialis), Steenbok (Raphicerus campestris), Greater kudu (Tragelaphus strepsiceros), Giraffe (Giraffa camelopardus), Burchells zebra (Equus burchelli), Blue wildebeest (Connocheatus taurinus), black-backed jackal (Canis mesomelas), Brown hyaena (Hyaena brunnea), Spotted hyaena (Crocuta crocuta), Lion (Panthera leo), Cheetah (Acinonyx jubatus), Painted hunting dog (Lycaon pictus) and even African bush elephant (Loxodonta africana) along the Boteti River. The Nxai Pan has a sizeable Springbok population and is one of the few places where Springbok and Impala cohabit. These two antelope are normally separated by habitat preference, but the Acacia savanna surrounding Nxai Pan provides the impala with a suitable habitat while the grass covered pan mimics the desert conditions preferred by Springbok.
- A. Campbell. 1990. The nature of Botswana: a guide to conservation and development. IUCN, Harare, Zimbabwe. ISBN: 2880329345
- C.MIchael Hogan & World Wildlife Fund. 2015. Zambezian halophytics. Encyclopedia of Earth. National Council for Science and Environment. Washington DC
Southern Africa Bushveld
Garman's toad (Amietophrynus garmani) is found in the Southern African bushveld, among other ecoregions. The Southern Africa bushveld is an element of the vast savannas that cover much of southern Africa. There is low endemism in this ecoregion for both flora or fauna, but the charismatic large mammals and rich birdlife characteristic of African savannas are in evidence. The rugged Waterberg Mountains contain the highest levels of species richness and endemism in the region, and are noted for their reptilian endemism. The ecoregion occurs on an extensive, undulating interior plateau, which lies at an elevation between 700 metres (m) to 1100 m. The soils of this plateau are chiefly coarse, sandy and shallow, overlying granite, quartzite, sandstone or shale. The most distinctive topographical feature of the ecoregion is the rugged and rocky Waterberg Mountains, which rise up from the plateau to an elevation of between 1200 m to 1500 m.http://www.eoearth.org/view/article/51cbeeed7896bb431f69b38d/554565bb0cf24df5070a17ee/?topic=51cbfc79f702fc2ba8129ee0
The ecoregion amphibian associates of the Southern African bushveld are: Savanna ridged frog (Ptychadena anchietae); Angola frog (Rana angolensis); African gray treefrog (Chiromantis xerampelina); Senegal running frog (Kassina senegalensis); Striped stream frog (Strongylopus fasciatus); African clawed frog (Xenopus laevis); African split-skin toad (Schismaderma carens); Uzungwe grassland frog (Ptychadena uzungwensis); African ornate frog (Hildebrandtia ornata); Mababe river frog (Phrynobatrachus mababiensis); Marbled sand frog (Tomopterna marmorata); Marbled snout burrower (Hemisus marmoratus); Knocking sand frog (Tomopterna krugerensis), which is found broadly in southern Africa; and the Transvaal short-headed frog (Breviceps adspersus); Mozambique ridged frog (Ptychadena mossambica); Lukula grassland frog (Ptychadena taenioscelis); Horseshoe forest treefrog (Leptopelis bocagii); South African snake-necked frog (Phrynomantis bifasciatus); Boettger's dainty frog (Cacosternum boettgeri); Natal ghost frog (Heleophryne natalensis); Cryptic sandfrog (Tomopterna cryptotis); Mozambique rain frog (Breviceps mossambicus); Long reed frog (Hyperolius nasutus); Muller's clawed frog (Xenopus muelleri); Common reed frog (Hyperolius viridiflavus); Gray's stream frog (Strongylopus grayii); Natal puddle frog (Phrynobatrachus natalensis); Painted reed frog (Hyperolius marmoratus); Garman's toad (Amietophrynus garmani); Gutteral toad (Amietophrynus gutturalis); Transvaal dwarf toad (Poyntonophrynus fenoulheti); and the Flat-back toad (Amietophrynus maculatus).
Example reptilian associates within this ecoregion are: Bibron's worm snake (Typhlops bibronii); Vine snake (Thelotornis capensis); Black mamba (Dendroaspis polylepis); Angola garter snake (Elapsoidea semiannulata); Annobon lidless skink (Panaspis annobonensis); Bark snake (Hemirhagerrhis nototaenia); Bell's hingeback tortoise (Kinixys belliana); Blue throated agama (Acanthocercus atricollis); Blunt-tailed worm lizard (Dalophia pistillum); Bradfield's dwarf gecko (Lygodactylus bradfieldi); the endemic gecko Broadley's rock gecko (Afroedura broadleyi); and the endemic lizards Platysaurus minor and Platysaurus monotropis.
Some of the many mammalian taxa found within the Southern African bushveld are: Burchell's zebra (Equus quagga burchelli); Hippopotamus (Hippopotamus amphibius), a herbivore classified as Vulnerable; Cheetah (Acinonyx jubatus), a carnivore classified as Vulnerable; the Near Threatened White Rhinoceros (Ceratotherium simum); Commerson's roundleaf bat (Tomopterna cryptotis), classified as Near Threatened; Spotted hyena (Crocuta crocuta); and the Mauritian tomb bat (Taphozous mauritianus).
There are numerous avian species found in this ecoregion, a few examples being: the Near Threatened Red footed falcon (Falco vespertinus); Kori bustard (Ardeotis kori); Long-crested eagle (Lophaetus occipitalis); Olive bee eater (Merops superciliosus); Marabou stork (Leptoptilos crumeniferus); Martial eagle (Polemaetus bellicosus); and the Pink-backed pelican (Pelecanus rufescens).
Habitat and Ecology
Movements and dispersal
Evolution and Systematics
The beak of the pink-backed pelican is usd to scoop and drain water during feeding via an extendable throat pouch.
"The pink-backed pelican uses its distendable throat pouch as a fishing net, scooping fish and crustaceans from the water as it swims. When it opens its bill underwater, the sudden inflow of water carries the prey in with it. Then the pelican raises its head to drain out the water before swallowing the prey." (Foy and Oxford Scientific Films 1982:157)
Learn more about this functional adaptation.
IUCN Red List Assessment
Red List Category
Red List Criteria
Status in Egypt
Regular passage visitor and winter visitor.
The pink-backed pelican (Pelecanus rufescens) is a bird of the pelican family. It is a resident breeder in swamps and shallow lakes of Africa, southern Arabia, southern India and is apparently extirpated in Madagascar.
This is a relatively small pelican though by no means a small bird. Length is from 125 to 155 cm (49 to 61 in), wingspan is 2.15–2.9 m (7.1–9.5 ft) and body mass if from 4 to 7 kg (8.8 to 15.4 lb). The bill is 30 to 38 cm (12 to 15 in) in length. The plumage is grey and white, with a pinkish hue on the back occasionally apparent (never in the deep pink of a flamingo). The top of the bill is yellow and the pouch is usually greyish. Breeding adults have long feather plumes on the head.
It shares habitat with the great white pelican which is generally larger and has white instead of greyish plumage.
Habitat and breeding
The pink-backed pelican is found in a range of aquatic habitats, but prefers quiet backwaters with shallow water, avoiding steep, vegetated lake banks. It prefers for freshwater lakes, swamps, large slow-flowing rivers, and seasonal pools but also frequents reservoirs, seasonally flooded land and flood-plains near river mouths. It may occur on alkaline and saline lakes and lagoons, and can sometimes be found along the coast in bays and estuaries (although seldom on open seashore). The species tends to roost and breed in trees (e.g. mangroves), but will also roost on sandy islands, cliffs, coral reefs and sand-dunes.
Nesting trees have many nests built close together, these nests are re-used every year until often the trees collapse although the birds will normally remain in the area. The species nests colonially in trees, reeds or low bushes along waterfronts as well as (less often) on the ground on sandy islands and in mangroves.
The nest is a large heap of sticks and may be 10–50 m above the ground. The female lays two to three large white eggs and later the chicks feed by plunging their heads deep into the adult’s pouch and taking the partially digested regurgitated fish.
Food is usually fish (of any size up to 450 g, usually in the 80-290 g range) and amphibians, and is usually obtained by fishing in groups. Among the fish preyed upon are cichlids like Haplochromis and Tilapia.