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Overview

Brief Summary

Biology

The diet is broad, incorporating seeds, roots, berries, leaves, and insects (4). Males mate with more than one female; females undertake the duties of nesting and rearing chicks alone (4). During late April, between 7 and 15 eggs are laid in a grass-lined hollow on the ground (4). The chicks hatch between 23 and 27 days later, and become independent after 12-14 days (4). Pheasants roost in trees (2), and form flocks in winter when feeding, in which hierarchies develop amongst the females (5)
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Comprehensive Description

Description

The pheasant was introduced to Britain by the Romans and Normans (4); further introductions of various races (or 'subspecies') have been made since (7), and it is now our commonest gamebird (5). As the different races have since interbred, adult plumage is extremely variable (4). Adult males are attractive and unmistakable, with a long tail, bright red wattles around the eyes (4), a chestnut coloured body, and an iridescent green or bluish head, which is often separated from the body by a white collar (4). Females are paler in colour, with spots and streaks, which provide good camouflage (4).
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Distribution

Global Range: Native to Asia. Introduced and established in most of Europe, New Zealand, and North America; also on all main islands of Hawaii. In North America from southern Canada south locally to California, Utah, southern New Mexico, southeastern Texas, northwestern Oklahoma, southern Illinois, Pennsylvania, New Jersey, and Maryland. Introduced on Guam but did not become established (Reichel and Glass 1991).

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Geographic Range

Phasianus colchicus is a non-migratory species native to Eurasia. The native range extends from the Caspian Sea, east across central Asia to China, and includes Korea, Japan, and former Burma. It was introduced to Europe, North America, New Zealand, Australia, and Hawaii. In North America, Phasianus colchicus populations have been established on mid-latitude agricultural lands from southern Canada to Utah, California to New England states, and south to Virginia.

Biogeographic Regions: nearctic (Introduced ); palearctic (Native ); australian (Introduced )

  • Federation of Alberta Naturalists, 2007. The Atlas of Breeding Birds of Alberta: a second look. Altona, Manitoba, Canada: Federation of Alberta Naturalists.
  • Giudice, J., J. Ratti. 2001. "The Birds of North America Online" (On-line). Ring-necked Pheasant. Accessed September 14, 2009 at http://bna.birds.cornell.edu/bna/species/572/articles/introduction.
  • Pratt, D., P. Bruner, D. Berrett. 1944. The Birds of Hawaii and the Tropical Pacific. Princeton, New Jersey: Princeton University Press.
  • Whitfield, P. 1984. Pheasants. Pp. 230-231 in P Whitfield, J Johnson, eds. Macmillan Illustrated Animal Encyclopedia. New York, NY: Macmillan Publishing Company.
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occurs (regularly, as a native taxon) in multiple nations

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National Distribution

Canada

Origin: Exotic

Regularity: Regularly occurring

Currently: Present

Confidence: Confident

Type of Residency: Year-round

United States

Origin: Exotic

Regularity: Regularly occurring

Currently: Present

Confidence: Confident

Type of Residency: Year-round

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Range

Pheasant shooting became popular in the 17th and 18th Centuries; large numbers of captive-reared birds are released each summer to supply this activity and supplement the population (4). The pheasant is now widely distributed throughout Britain, but is absent from the West Highlands and the islands of Scotland, and from some areas of the uplands in England and Wales (5). It is known throughout western Europe, central Asia, China, Korea and southeastern Siberia, and has been introduced to many other areas (6).
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Physical Description

Morphology

Physical Description

Common pheasants are medium-sized birds with deep, pear-shaped bodies, small heads and long, thin tails. They are sexually dimorphic, with males being more colorful and larger than females. Males have spectacular, multi-colored plumage with long, pointed, barred tails and fleshy red eye patches. Their heads range in color from glossy dark green to iridescent purple. Many subspecies have a distinctive white collar around their neck which gives them their ‘ring-necked’ name. Female Phasianus colchicus are less colorful. They have buff brown, mottled plumage and, like males, have long pointed tails, although they are shorter than those of males.

There are two major groupings of subspecies within Phasianus colchicus. The colchicus group, or ring-necked pheasants, are native to mainland Eurasia. They are barred, with coppery red or yellow on their mantle and underparts, and have the prominent neck ring. Thirty-one subspecies are listed under this grouping. The other grouping of subspecies is the versicolor group, which lacks the neck ring and has green on their neck, breast, and upper belly. This group is native to Japan and was introduced to Hawaii. There are three subspecies in the versicolor group.

Average mass: 1263 g.

Range length: 42.5 to 53.6 cm.

Range wingspan: 23.5 to 25.8 cm.

Other Physical Features: endothermic ; homoiothermic; bilateral symmetry

Sexual Dimorphism: male larger; sexes colored or patterned differently; male more colorful; ornamentation

  • Delacour, J. 1977. The Pheasants of the World. Surrey, England: Spur Publications.
  • Greenberg, R. 2002. Ring-necked Pheasant. Pp. 185 in J Greenberg, J Hamilton, eds. Birds of Canada. Kyodo, Singapore: Dorling Kindersley.
  • Johnsgard, P. 1975. North American Game Birds of Upland and Shoreline. Lincoln, Nebraska: University of Nebraska Press.
  • Johnsgard, P. 1986. The Pheasants of the World. Oxford, New York: Oxford University Press.
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Size

Length: 84 cm

Weight: 1317 grams

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Ecology

Habitat

Habitat and Ecology

Systems
  • Terrestrial
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Habitat

Common pheasants occupy grassland and farmland habitats. They prefer relatively open cover, such as grass and stubble fields and are found in habitats with grass, ditches, hedges, marshes, and tree stands or bushes for cover. They are generalists occupying a wide range of habitat types except areas with dense rainforest, alpine forests, or very dry places. This flexibility is exemplified in their successful introduction to tropical habitats in Hawaii where only heavy precipitation and high altitudes pose the greatest habitat limitation.

Open water is not a requirement for Phasianus colchicus, but most populations are found where water is present. In drier habitats, common pheasants obtain water from dew, insects, and succulent vegetation.

Common pheasants occupy agricultural areas but the movement towards increasingly large agricultural operations is detrimental to habitat. Land-use transitions to larger operations include a loss of field-edge habitat (fewer fencerows), removal of bushes, burning of marshes, a trend towards monoculture, suburban sprawl, and commercial development. This habitat degradation leads to reduction of cover habitat and fewer small bodies of water for Phasianus colchicus.

Range elevation: 0 to 3353 m.

Habitat Regions: temperate ; tropical ; terrestrial

Terrestrial Biomes: savanna or grassland

Wetlands: marsh

Other Habitat Features: agricultural

  • Schwartz, C., E. Shwartz. 1951. An Ecological Reconnaissance of the Pheasants in Hawaii. The Auk, 68/3: 281-314.
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Comments: Open country (especially cultivated areas, scrubby wastes, open woodland and edges of woods), grassy steppe, desert oases, riverside thickets, swamps and open mountain forest (AOU 1983). Winter shelter includes bushes and trees along streams, shelterbelts, and fencerows. Usually nests in fields, brushy edges, or pastures, also along road rights-of-way. Nest is shallow depression scratched out by female.

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Habitat

Typically prefers wooded agricultural lowland (5), but may also occur in gardens, parks and marshes, their preferred habitats in Asia (4).
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Migration

Non-Migrant: Yes. At least some populations of this species do not make significant seasonal migrations. Juvenile dispersal is not considered a migration.

Locally Migrant: No. No populations of this species make local extended movements (generally less than 200 km) at particular times of the year (e.g., to breeding or wintering grounds, to hibernation sites).

Locally Migrant: No. No populations of this species make annual migrations of over 200 km.

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Trophic Strategy

Food Habits

Common pheasants are dietary generalists, eating a wide variety of plant matter, such as grain, seeds, shoots, and berries, as well as insects and small invertebrates. Common pheasants are mostly ground dwelling and scratch for food in the undergrowth with their bill. They usually forage in the early morning and evening. Important agricultural crops eaten by common pheasants are corn (Zea mays), wheat (Triticum), barley (Hordeum vulgare), and flax (Lineum) Weed seeds they eat in North America are foxtail (Setaria lutescens), ragweed (Ambrosia) and sunflower seeds (Helianthus annus). Wild grape (Vitis), apples (Malus), and blackberries (Rubus) are some fruits eaten. They also eat grasshoppers (Orthoptera), caterpillars (Lepidoptera), crickets (Gryllidae), and snails (Gastropoda).

Animal Foods: insects; terrestrial non-insect arthropods

Plant Foods: seeds, grains, and nuts; fruit

Primary Diet: omnivore

  • Dale, F., J. Lauckhart, J. McKean. 1956. Pheasants in North America. Harrisburg, Pennsylvania: The Stackpole Company.
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Comments: Eats waste corn, wheat, barley, oats, buckwheat, and seeds of ragweed, burdocks, etc.; also berries and pine seeds. In the spring eats green vegetation. Also eats some insects, mice, and snails (Terres 1980). Able to scratch through snow to reach food on the ground surface.

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Associations

Ecosystem Roles

Common pheasants play a role as prey for larger carnivores and as an insectivore, helping to control insect populations. They may also disperse seeds through their seed predation. They may negatively affect greater prairie chickens (Tympanuchus cupido) and gray partridges (Perdix perdix) through nest parasitism, habitat competition, transmission of disease, and aggressive behaviour. A study in Kansas reported nest parasitism of lesser prairie chickens by common pheasants. This rate of nest parasitism appeared to be density-dependent, increasing as nest site availability decreased. Other studies have investigated the negative impact of common pheasants on greater prairie chickens through nest parasitism. Pheasant eggs hatch earlier than prairie chicken eggs. The rate of embryo mortality or nest abandonment increases in parasitized prairie-chicken nests.

The release of common pheasants into woodland areas for game shooting is common. One study in Britain looked into the impact of this practice. They found that there was a neutral or positive impact of common pheasants on vegetation and bird communities. However, it is important to note that this study was done in pheasant-managed woodland areas and this management practice may have been more beneficial than the presence of pheasants themselves.

Ecosystem Impact: disperses seeds

Commensal/Parasitic Species:

  • Aldous, E., D. Alexander. 2008. Newcastle disease in pheasants (Phasianus colchicus): A review. The Veterinary Journal, 175/2: 181-185.
  • Draycott, R., A. Hoodless, R. Sage. 2008. Effects of pheasant management on vegetation and birds in lowland woodlands. Journal of Applied Ecology, 45/1: 334-341.
  • Hagen, C., B. Jamison, R. Robel, R. Applegate. 2002. Ring-necked Pheasant Parasitism of Lesser Prairie Chicken Nests in Kansas. The Wilson Bulletin, 114/4: 522-524.
  • Westemeier, R., J. Buhnerkempe, W. Edwards, J. Brawn, S. Simpson. 1998. Parasitism of Greater Prairie-Chicken Nests by Ring-necked Pheasants. The Journal of Wildlife Management, 62/3: 854-863.
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Predation

Adult common pheasants may be preyed on either while on the ground or in flight. Some of their behavioral responses to danger include retreating into cover or hiding. They also may fly, crouch, or run. Hens facing a predator may display a broken wing in an attempt to draw their predator away from their nest or they may just try to sit very still. When chicks in a brood are preyed on, often more than one is taken at a time. Exposure to extreme weather is also attributed to chick mortality. Game hunting by humans is a significant predation concern for male pheasants in some areas. Common pheasants are particularly vulnerable to predation during nesting. Studies have shown that control of nest predators, particularly red foxes, can be a significant pheasant conservation tool. Additionally, increased pheasant predation rates are linked closely to increased rates of habitat destruction. This may be because habitat degradation renders prey more vulnerable to predators.  Studies have also been conducted to determine whether certain subspecies have higher survival rates in specific habitats. One particular study focused on the Sichuan subspecies of common pheasants, which nest in woody cover, a trait which makes them less susceptible to agricultural land degradation. However, this study found that Sichuan hens had no survival advantage over hens of other subspecies. Much information on predation in common pheasants is known from North American populations, where they are an important game species.

Known Predators:

Anti-predator Adaptations: cryptic

  • Draycott, R., A. Hoodless, M. Woodburn, R. Sage. 2008. Nest predation of Common Pheasants Phasianus colchicus. Ibis, 150/1: 37-44.
  • Shipley, K., D. Scott. 2006. Survival and Nesting Habitat use by Sichuan and Ring-necked Pheasants Released in Ohio. Ohio Journal of Science, 106/3: 78-85.
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Known predators

Phasianus colchicus (ring-necked pheasent) is prey of:
Buteo swainsoni

Based on studies in:
USA: California, Cabrillo Point (Grassland)

This list may not be complete but is based on published studies.
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Associations

Animal / dung saprobe
apothecium of Ascobolus crenulatus is saprobic in/on dung or excretions of dung of Phasianus colchicus

In Great Britain and/or Ireland:
Animal / parasite / ectoparasite / blood sucker
Ixodes ricinus sucks the blood of Phasianus colchicus
Other: major host/prey

Animal / predator
adult of Phasianus colchicus is predator of adult of Timarcha tenebricosa

Animal / dung saprobe
immersed, neck protruding perithecium of Podospora curvicolla is saprobic in/on dung or excretions of dung of Phasianus colchicus
Other: minor host/prey

Animal / dung saprobe
partly immersed perithecium of Podospora pleiospora is saprobic in/on dung or excretions of dung of Phasianus colchicus

Animal / dung saprobe
scattered or gregarious, superficial, sessile apothecium of Saccobolus dilutellus is saprobic in/on dung or excretions of dung of Phasianus colchicus

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General Ecology

Usually ranges over no more than approximately 2-3 km (Terres 1980). In fall family groups may join and form flocks of 30-40 birds. Flocks break up in spring.

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Life History and Behavior

Behavior

Communication and Perception

When alarmed, common pheasants make distinctive hoarse croaking notes. In males, this is a loud, piercing, double squawk ko-ork kok, with a sharp staccato on the last syllable. This crowing call is also made when males are establishing their territory. In agricultural areas, males may be heard crowing at dusk, dawn, and during the mating season. This call is very similar to the familiar call of a rooster and can travel up to a mile. Female calls tend to be more subtle and less likely to be audible.

Communication Channels: visual ; acoustic

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Life Expectancy

Lifespan/Longevity

Chick survival is influenced by hatch date, mass at birth, and habitat type. Many young don’t live beyond autumn. Annual survival rate of adult females is 21 to 46%, while it is only 7% for males. In some areas the reduced survival rate of males can be accounted for by the hunting of male common pheasants by humans. Nearly all wild birds die by age three. Adult mortality is caused by predation, agricultural activities, exposure to pesticides and toxins, and accidents with motorized vehicles.

Range lifespan

Status: captivity:
11 to 18 years.

Average lifespan

Status: wild:
3 years.

  • Martin, P., D. Johnson, D. Forsyth. 1996. Effects of grasshopper-control insecticides on survival and brain acetylcholinesterase of pheasant (Phasianus colchicus) chicks. Environmental Toxicology and Chemisty, 15/4: 518-524.
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Lifespan, longevity, and ageing

Maximum longevity: 27 years (captivity) Observations: In the wild, mortality is high and only 5% of animals survive up to age 3 (http://www.fs.fed.us/database/feis/).
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Reproduction

Reproduction

Common pheasants are polygynous, with a single male having a harem of several females. Common pheasants breed seasonally. In early spring (mid-March to early June) males establish breeding or crowing territories. These territories are relative in terms of other males’ territories and do not necessarily have definitive boundaries. On the other hand, females are not territorial. Within their breeding harem, they may show a dominance hierarchy. These harems last through the courting and nesting period and may have 2 to 18 females. Each female typically has a seasonally monogamous relationship with one territorial male.  In early spring, males establish a harem by crowing and wing-whirring displays. Crowing is the distinctive, loud korrk-kok call of males which they use to maintain their territory. This may be preceded by an almost inaudible wing-flap, after which the male may perform a brief but vigorous wing-whirring. Physical interactions between competing males may include flying at each other breast-to-breast, biting wattles, or high leaps with kicks toward the other’s bill. Males who establish breeding territories earlier in the season tend to be dominant over males establishing territory later.  Mate selection by females is dependent on a few factors. Female common pheasants tend to choose dominant males who can, for example, offer protection. Studies have found that females prefer long tails in males and that the length of ear tufts and presence of black points on the wattle also influences female choice. The general brightness of a male's plumage is not a factor, perhaps because brightness is not correlated to testosterone levels or dominant behaviors in male common pheasants.

Males have different courtship displays which elicit different responses in females. One study found that feeding rituals in males attracted female common pheasants, while lateral display courtship behaviors in males aroused females for copulation. In a lateral display, the male approaches the female, crossing slowly in a semicircle in front of her with his head low, the nearer wing drooped and his wattle erect. This lateral display often precedes copulation but later in the season a male may simply pursue and attempt to mount a female.

Mating System: polygynous

Nesting begins just before females start to lay eggs. The female will scrape a shallow depression in the ground in a well covered area, lining it lightly with readily available plant material. She will typically lay one egg a day until 7 to 15 eggs have been laid. Larger clutches of eggs arise when two or more hens lay in the same nest. The female will remain close to the nest, incubating the eggs for most of the day, leaving in the morning and evening to feed. Chicks are precocial at hatching, completely covered with down and with their eyes open. They are able to immediately begin walking and following the hen to sources of food; they are largely self-feeding.

Breeding interval: Ring-necked pheasants breed once yearly.

Breeding season: The ring-necked pheasant breeding season extends from mid-March to June.

Range eggs per season: 7 to 15.

Average eggs per season: 10.

Range time to hatching: 23 to 28 days.

Average time to hatching: 24 days.

Average birth mass: 18.5 g.

Range fledging age: 7 to 12 days.

Range time to independence: 70 to 80 days.

Average age at sexual or reproductive maturity (female): 1 years.

Average age at sexual or reproductive maturity (male): 1 years.

Key Reproductive Features: seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate)

Most parental investment in common pheasants is by females. After building her nest and laying the eggs, the female is responsible for incubating them. Incubation takes approximately 23 days after the final egg is laid. When the chicks hatch, they are cared for solely by the hen. They are precocial when they hatch, covered with down, eyes open, and legs developed. They are able to immediately begin following the hen to sources of food and the young chicks will feed themselves. The hen’s main role is to lead her chicks to food after hatching. By about 12 days, young are able to fly and typically remain with the hen for 70 to 80 days before becoming independent.

Parental Investment: precocial ; female parental care ; pre-fertilization (Provisioning, Protecting: Female); pre-hatching/birth (Provisioning: Female, Protecting: Female); pre-weaning/fledging (Protecting: Female)

  • Giudice, J., J. Ratti. 2001. "The Birds of North America Online" (On-line). Ring-necked Pheasant. Accessed September 14, 2009 at http://bna.birds.cornell.edu/bna/species/572/articles/introduction.
  • Greenberg, R. 2002. Ring-necked Pheasant. Pp. 185 in J Greenberg, J Hamilton, eds. Birds of Canada. Kyodo, Singapore: Dorling Kindersley.
  • Grzimek, B., G. Archibald, C. Berger. 2003. Fowls and pheasants (phasianidae). Pp. 433-439 in M Hutchins, W Bock, J Jackson, eds. Grzimek's Animal Life Encyclopedia, Vol. 8, 2nd Edition. Farmington Hills, MI: Gale Group.
  • Johnsgard, P. 1975. North American Game Birds of Upland and Shoreline. Lincoln, Nebraska: University of Nebraska Press.
  • Johnsgard, P. 1986. The Pheasants of the World. Oxford, New York: Oxford University Press.
  • Mateos, C., J. Carranza. 1999. Effects of male dominance and courtship display on female choice in the ring-necked pheasant. Behavioral Ecology and Sociology, 45/3-4: 235-244.
  • Mateos, C., J. Carranza. 1995. Female choice for morphological features of male ring-necked pheasants. Animal Behaviour, 49/3: 737-748.
  • Whitfield, P. 1984. Pheasants. Pp. 230-231 in P Whitfield, J Johnson, eds. Macmillan Illustrated Animal Encyclopedia. New York, NY: Macmillan Publishing Company.
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Usually 10-12, sometimes 5-23 eggs are laid in one nest. Two hens may lay eggs in same nest. Incubation lasts 23-25 days. Precocial young are tended by female.

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Molecular Biology and Genetics

Molecular Biology

Barcode data: Phasianus colchicus

The following is a representative barcode sequence, the centroid of all available sequences for this species. 

 
There are 14 barcode sequences available from BOLD and GenBank.  Below is a sequence of the barcode region Cytochrome oxidase subunit 1 (COI or COX1) from a member of the species.  See the BOLD taxonomy browser for more complete information about this specimen and other sequences.
 
KBPBU115-06|UWBM 46494|Phasianus colchicus| ------------------------------------------CTTTATCTAATTTTCGGCACATGAGCGGGCATAGTTGGCACAGCACTT---AGCCTACTAATTCGCGCAGAACTAGGACAACCAGGAACACTTTTGGGAGAT---GACCAAATTTATAATGTAATCGTCACAGCCCATGCCTTCGTTATAATCTTTTTCATAGTTATACCTATCATGATTGGGGGCTTCGGAAATTGACTAGTCCCTCTTATA---ATCGGTGCTCCAGACATAGCATTCCCACGTATAAACAACATAAGCTTCTGACTTCTCCCCCCCTCCTTTCTTCTCCTGCTAGCCTCCTCAACCGTAGAAGCTGGAGCCGGTACCGGATGAACTGTTTACCCCCCCTTAGCTGGAAACCTTGCTCACGCCGGTGCATCAGTAGATCTG---GCCATTTTCTCCCTCCATCTCGCAGGTGTGTCATCTATTTTAGGGGCCATTAACTTCATCACTACCATCATCAACATAAAACCCCCTGCATTATCACAATACCAAACACCACTATTCGTCTGATCCGTCCTTATTACCGCCATTCTCCTACTACTTTCTTTACCCGTTCTAGCCGCT---GGGATTACTATACTCCTTACTGACCGCAACCTCAACACCACATTCTTTGATCCTGCAGGAGGGGGAGATCCAATCCTCTATCAACACCTATTTTGATTCTTCGGCCATCCCGAAGTTTACATCCTCATCCTC------------------------------------------------------------------------------------------------------------------------------------------------------- 
-- end --

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Statistics of barcoding coverage: Phasianus colchicus

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 14
Species: 19
Species With Barcodes: 1

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Conservation

Conservation Status

IUCN Red List Assessment


Red List Category
LC
Least Concern

Red List Criteria

Version
3.1

Year Assessed
2009

Assessor/s
BirdLife International

Reviewer/s
Bird, J., Butchart, S.

Contributor/s

Justification
This species has an extremely large range, and hence does not approach the thresholds for Vulnerable under the range size criterion (Extent of Occurrence <20,000 km2 combined with a declining or fluctuating range size, habitat extent/quality, or population size and a small number of locations or severe fragmentation). Despite the fact that the population trend appears to be decreasing, the decline is not believed to be sufficiently rapid to approach the thresholds for Vulnerable under the population trend criterion (>30% decline over ten years or three generations). The population size is extremely large, and hence does not approach the thresholds for Vulnerable under the population size criterion (<10,000 mature individuals with a continuing decline estimated to be >10% in ten years or three generations, or with a specified population structure). For these reasons the species is evaluated as Least Concern.

History
  • 2008
    Least Concern
  • 2004
    Least Concern
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Conservation Status

Common pheasants are widely distributed and their conservation status is of least concern.

US Migratory Bird Act: no special status

US Federal List: no special status

CITES: no special status

State of Michigan List: no special status

IUCN Red List of Threatened Species: least concern

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National NatureServe Conservation Status

Canada

Rounded National Status Rank: NNA - Not Applicable

United States

Rounded National Status Rank: NNA - Not Applicable

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NatureServe Conservation Status

Rounded Global Status Rank: G5 - Secure

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Status

Introduced (9). Covered by Game Acts which give protection in the close season and allow it to be shot from 1st October to 1st February (10).
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Population

Population
In Europe, the breeding population is estimated to number 3400000-4700000 breeding pairs, equating to 10200000-14100000 individuals (BirdLife International 2004). Europe forms 5-24% of the global range, so a very preliminary estimate of the global population size is 45000000-300000000 individuals, although further validation of this estimate is needed.
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Threats

Threats

Not currently threatened.
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Management

Conservation

The Game Conservancy Trust is currently researching the consequences of releasing large numbers of captive-bred pheasants into small areas (8).
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Relevance to Humans and Ecosystems

Benefits

Economic Importance for Humans: Negative

Common pheasants are highly susceptible to Newcastle disease, a significant disease in birds because of high mortality rates in those affected. Disease outbreaks have economic implications including trade embargos and restrictions of poultry sales in areas of outbreak. Common pheasants can carry Newcastle disease and spread it to other wild and domestic birds, which can be potentially negative to humans.

Negative Impacts: causes or carries domestic animal disease

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Economic Importance for Humans: Positive

The predominant benefit of Phasianus colchicus to humans is as an upland game bird.

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Wikipedia

Common Pheasant

The Common Pheasant (Phasianus colchicus), is a bird in the pheasant family (Phasianidae). It is native to Georgia (Ancient Colchis) and has been widely introduced elsewhere as a game bird. In parts of its range, namely in places where none of its relatives occur such as in Europe (where it is naturalized), it is simply known as the "pheasant". Ring-necked Pheasant is both the name used for the species as a whole in North America and also the collective name for a number of subspecies and their intergrades which have white neck rings.

The word pheasant is derived from the ancient town of Phasis, the predecessor of modern Poti in Western Georgia). See below for details.

It is a well-known gamebird, among those of more than regional importance perhaps the most widespread and ancient one in the whole world. The Common Pheasant is one of the world's most hunted birds;[1] it has been introduced for that purpose to many regions, and is also common on game farms where it is commercially bred. Ring-necked Pheasants in particular are commonly bred and were introduced to many parts of the world; the game farm stock, though no distinct breeds have been developed yet, can be considered semi-domesticated. The Ring-necked Pheasant is the state bird of South Dakota, one of only three US state birds that is not a species native to the United States.

The Green Pheasant (P. versicolor) of Japan is sometimes placed as subspecies within the Common Pheasant. Though the species produce fertile hybrids wherever they coexist, this is simply a typical feature among fowl (Galloanseres), in which postzygotic isolating mechanisms are slight compared to most other birds. The species apparently have somewhat different ecological requirements and at least in its typical habitat the Green outcompetes the Common Pheasant; its introduction to Japan has therefore largely failed.

Contents

Description

There are many colour forms of the male Common Pheasant, ranging in colour from nearly white to almost black in some melanistic examples. These are due to captive breeding and hybridization between subspecies and with the Green Pheasant, reinforced by continual releases of stock from varying sources to the wild. For example, the "Ring-necked Pheasants" common in Europe, North America and Australia do not pertain to any specific taxon, they rather represent a stereotyped hybrid swarm.[2] Body weight can range from 0.5 to 3 kg (1.1-6.6 lb), with males averaging 1.2 kg (2.6 lb) and females averaging 0.9 kg (2 lb).[3][4]

The adult male Common Pheasant of the nominate subspecies Phasianus colchicus colchicus is 60–89 cm (24–35 in) in length with a long brown streaked black tail, accounting for almost 50 cm (20 in) of the total length. The body plumage is barred bright gold and brown plumage with green, purple and white markings. The head is bottle green with a small crest and distinctive red wattle. P. c. colchicus and some other races lack a white neck ring.

Flavistic hen

The female (hen) is much less showy, with a duller mottled brown plumage all over and measuring 50–63 cm (20–25 in) long including a tail of around 20 cm (8 in). Juvenile birds have the appearance of the female with a shorter tail until young males begin to grow characteristic bright feathers on the breast, head and back at about 10 weeks after hatching.

The Green Pheasant (P. versicolor) is very similar, and hybridizaton makes individual birds' identities often difficult to determine. Green Pheasant males are shorter-tailed on average and have a darker plumage that is uniformly bottle-green on the breast and belly; they always lack a neck ring. Their females are darker, with many black dots on the breast and belly.

In addition, various color mutations are commonly encountered, mainly melanistic (black) and flavistic (isabelline or fawn) specimens. The former are rather common in some areas and are named Tenebrosus Pheasant (P. colchicus var. tenebrosus).

Taxonomy and systematics

Photo of a bird
male Common Pheasant
Hybrid male in Europe, intermediate between Mongolian Ringneck and Caucasus group phenotype

This species was first scientifically described by Linnaeus in his Systema naturae in 1758 under its current scientific name. The Common Pheasant is distinct enough from any other species known to Linnaeus for a laconic [Phasianus] rufus, capîte caeruleo – "a red pheasant with blue head" – to serve as entirely sufficient description. Moreover, the bird had been extensively discussed before Linnaeus established binomial nomenclature. His sources are the Ornithologia of Ulisse Aldrovandi,[5] Giovanni Pietro Olina's Uccelliera,[6] John Ray's Synopsis methodica Avium & Piscium,[7] and A natural history of the birds by Eleazar Albin.[8] Therein – essentially the bulk of the ornithology textbooks of his day – the species is simply named "the pheasant" in the books' respective languages. Whereas in other species, such as the Eastern Meadowlark (Sturnella magna), Linnaeus felt it warranted to cite plumage details from his sources, in the Common Pheasant's case he simply referred to the reason of the bird's fame: principum mensis dicatur[verification needed]. The type locality is given simply as "Africa, Asia".[9]

However, the bird does not occur in Africa, except perhaps in Linnaeus' time in Mediterranean coastal areas where they might have been introduced during the Roman Empire. The type locality was later fixed to the Rioni River – known as Phasis to the Ancient Greeks – where the westernmost population occurs. These birds, until the modern era, constituted the bulk of the introduced stock in Europe; the birds described by Linnaeus' sources, though typically belonging to such early introductions, would certainly have more alleles in common with the transcaucasian population than with others. The scientific name is Latin for "Pheasant from Colchis", colchicus referring to the west of modern-day Georgia;[10] the Ancient Greek term corresponding to the English "pheasant" is Phasianos ornis (Φασιανὸς ὂρνις), "bird of the river Phasis".[11] Although Linnaeus included many Galliformes in his genus Phasianius – such as the domestic chicken and its wild ancestor the Red Junglefowl, nowadays Gallus gallus –, today only the Common and the Green Pheasant are placed in this genus. As the latter was not known to Linnaeus in 1758, the Common Pheasant is naturally the type species of Phasianus.

In the USA, Common Pheasants are widely known as "Ring-necked Pheasants". More colloquial North American names include "chinks" or, in Montana, "phezzens".[12] In China, meanwhile, the species is properly called zhi ji (雉鸡) – "pheasant-fowl" –, essentially implying the same as the English name "Common Pheasant". Like elsewhere, P. colchicus is such a familiar bird in China that it is usually just referred to as shan ji (山雞), "mountain chicken",[13] a Chinese term for pheasants in general.

As of 2005, it had the smallest known genome of all living amniotes, only 0.97 pg (970 million base pairs), however Black-chinned Hummingbird is now currently known to have the smallest.[14]

Subspecies

Caucasus Pheasant-type male
Mongolian Ringneck-type male
Chinese Ringneck-type male (note grey rump) with very pale female

There are about 30 subspecies in five (sometimes six) groups.[15] These can be identified according to the male plumage, namely presence or absence of a white neck-ring and the color of the uppertail (rump) and wing coverts. As noted above, introduced population in our time mix the alleles of various races in various amounts, differing according to the original stock used for introductions and what natural selection according to climate and habitat has made of that.

Sometimes this species is split into the Central Asian Common and the East Asian Ring-necked Pheasants, roughly separated by the arid and high mountainous regions of Turkestan. However, while the western and eastern populations probably were entirely separate during the Zyryanka glaciation when deserts were more extensive,[16] this separation was not long enough for actual speciation to occur. Today, the largest variety of color patterns is found where the western and eastern populations mix, as is to be expected. Females usually cannot be identified even to subspecies group with certainty.

The subspecies groups, going from west to east, are:

Caucasus to W Turkestan.
No neck ring. Wing coverts buff to brown, uppertail coverts rusty to chestnut.
  • Phasianus colchicus chrysomelas/principalis group – White-winged Pheasants including Prince of Wales Pheasant (P. c. principalis)
Central Turkestan.
No or vestigial neck ring. Wing coverts white, uppertail coverts and general plumage hue bronze to brown.
NE Turkestan and adjacent Mongolia.
Broad neck ring. Wing coverts white, uppertail coverts hue rusty to chestnut, general plumage hue copper.
SE Turkestan around the Tarim Basin.
No or vestigial neck ring. Wing coverts buff to brown, uppertail coverts dark khaki to light olive.
Throughout China but widespread in the east, extending to northernmost Vietnam and Taiwan in the south and to the Strait of Tartary region in the north. Most pheasants in North America are of this group.
Usually broad neck ring. Wing coverts tan to light grey (almost white in some), uppertail coverts grey to powder blue with orange tips. Top of head light grey.

Ecology

A startled male makes a dash for cover

Common Pheasants are native to Asia, their original range extending from between the Black and Caspian Seas to Manchuria, Siberia, Korea, Mainland China and Taiwan. The birds are found in woodland, farmland, scrub and wetlands. In its natural habitat the Common Pheasant lives in grassland near water with small copses of trees. Extensively cleared farmland is marginal habitat that cannot maintain self-sustaining populations for long[17].[15]

Common Pheasants are gregarious birds and outside the breeding season form loose flocks. Wherever they are hunted they are always timid once they associate humans with danger, and will quickly retreat for safety after hearing the arrival of hunting parties in the area.

Chicks about 1 hour after hatching

While Common Pheasants are able short-distance fliers, they prefer to run. If startled however, they can suddenly burst upwards at great speed, with a distinctive "whirring" wing sound and often giving kok kok kok calls to alert conspecifics. Their flight speed is only 43-61 kilometres per hour (27 to 38 mph) when cruising but when chased they can fly up to 90 kilometres per hour (60 mph).

Common Pheasants feed solely on the ground but roost in sheltered trees at night. They eat a wide variety of animal and vegetable type-food, like fruit, seeds and leaves as well as a wide range of invertebrates, with small vertebrates like snakes, lizards, small mammals and birds occasionally taken.

The males are polygynous as is typical for many Phasianidae, and are often accompanied by a harem of several females.[18] Common Pheasants nest on the ground, producing a clutch of around ten eggs over a two-three week period in April to June. The incubation period is about 23–26 days. The chicks stay near the hen for several weeks after hatching but grow quickly, resembling adults by only 15 weeks of age.

As introduced species

Although a non-indigenous species, the Common Pheasant is farmed even in conservation areas, as seen here in Litovelské Pomoraví Protected Landscape Area in the Czech Republic.
Introduced birds sometimes forage in gardens
Collisions between pheasants and road vehicles are common in the UK[19]

Common Pheasants can now be found across the globe due to their readiness to breed in captivity and the fact they can naturalise in many climates. Pheasants were hunted in their natural range by Stone Age humans just like the grouse, partridges, junglefowls and perhaps peacocks that inhabited Europe at that time. At least since the Roman Empire the bird was extensively introduced in many places and has become a naturalized member at least of the European fauna. Introductions in the Southern Hemisphere have mostly failed, except where local Galliformes or their ecological equivalents are rare or absent.

The bird was naturalized in Great Britain around the 10th century AD, arguably earlier, by both the Romano-British[20] and the Normans, but became extirpated from most of the isles in the early 17th century. It was rediscovered as a gamebird in the 1830s after being ignored for many years. Since then it has been reared extensively by gamekeepers. Because around 30 million pheasants are released each year on shooting estates, it is widespread in distribution, although most released birds survive less than a year in the wild. The Game & Wildlife Conservation Trust is researching the breeding success of reared pheasants and trying to find ways to improve this breeding success to reduce the demand to release as many reared pheasants and increase the wild population. As the original Caucasian stock all but disappeared during the Early Modern era, most dark-winged ringless birds in the UK are actually descended from Chinese Ringneck and Green Pheasant hybrids[21] which were commonly used for rewilding.

Common Pheasants were introduced in North America in 1881,[22][23] and have become well established throughout much of the Rocky Mountain states (Colorado, Montana, Wyoming, etc.), the Midwest, the Plains states, as well as Canada and Mexico. In the southwest, they can even be seen south of the Rockies in Bosque del Apache National Wildlife Refuge 100 miles south of Albuquerque, New Mexico. It is now most common on the Great Plains. Common Pheasants have also been introduced to much of north-west Europe, the Hawaiian Islands, Chile, St Helena, Tasmania, New Zealand and Rottnest Island off Australia. It has also been unsuccessfully introduced to many other countries.

As gamebirds

An Field-bred English Cocker Spaniel has brought in the quarry

Common Pheasants are bred to be hunted and are shot in great numbers in Europe, especially the UK, where they are shot on the traditional formal "driven shoot" principles, whereby paying guns have birds driven over them by beaters, and on smaller "rough shoots". The open season in the UK is 1 October - 1 February, under the Game Act 1831. Generally they are shot by hunters employing gun dog to help find, flush, and retrieve shot birds. Retrievers, spaniels, and pointing breeds are used to hunt pheasants.

The doggerel "Up gets a guinea, bang goes a penny-halfpenny, and down comes a half a crown" reflects the expensive sport of nineteenth century driven shoots in Britain,[24] when pheasants were often shot for sport rather than as food. It was a popular royal pastime in Britain to shoot Common Pheasants. King George V shot over a thousand pheasants out of a total bag of 3937 over a six day period in December 1913 during a competition with a friend, however did not do enough to beat him.[21]

Common Pheasants are traditionally a target of small game poachers in the UK, but due to low value of pheasants in the modern day some have resorted to stealing chicks or poults from pens.[25] The Roald Dahl novel Danny the Champion of the World dealt with a poacher (and his son) who lived in the United Kingdom and illegally hunted Common Pheasants.

Just hatched, in an egg incubator

Pheasant farming is a common practice, and is sometimes done intensively. Birds are supplied both to hunting preserves/estates and restaurants, with smaller numbers being available for home cooks. Pheasant farms have some 10 million birds[citation needed] in the U.S. and 35 million in the United Kingdom.

For sale at Borough Market, London

The carcasses were often hung for a time to improve the meat by slight decomposition, as with most other game. Modern cookery generally uses moist roasting and farm-raised female birds. In the U.K. and U.S., game is making somewhat of a comeback in popular cooking, and more pheasants than ever are being sold in supermarkets there.[26] A major reason for this is consumer attitude shift from consumption of red meat to white.[27]

Pheasant hunting in North America

Most common pheasants bagged in the United States are wild-born feral pheasants; in some states[28] captive-reared and released birds make up much of the population[29]

Pheasant hunting is very popular in much of the U.S, especially in the Great Plains states where a mix of farmland and native grasslands create ideal habitat. South Dakota alone has an annual harvest of over a million birds a year by over 150,000 hunters.[30] Much of the North American hunting is done by groups of hunters with flushing dogs such as Labrador Retrievers and Springer Spaniels walking through fields and shooting the birds as they take flight. There are also many hunters who use Pointers such as English Setters or German Shorthairs to find and hold pheasants for hunters to flush and shoot.

Footnotes

  1. ^ Robertson (1997) pp.123-136
  2. ^ Sibley (2000): p.141
  3. ^ "Ring-necked Pheasant, Life History, All About Birds - Cornell Lab of Ornithology". Allaboutbirds.org. http://www.allaboutbirds.org/guide/Ring-necked_Pheasant/lifehistory. Retrieved 2011-04-25. 
  4. ^ "Ring-necked pheasant: Definition from". Answers.com. 2009-05-12. http://www.answers.com/topic/common-pheasant. Retrieved 2011-04-25. 
  5. ^ Aldrovandi (1600): pp.45-59
  6. ^ Olina (1622): p.49, plate 48
  7. ^ Ray (1713): p.56
  8. ^ Albin (1731): pp.24-26
  9. ^ Linnaeus (1758)
  10. ^ URB (2007)
  11. ^ Oxford English Dictionary Online (2008)
  12. ^ Proper (1990): pp.21-22
  13. ^ E.g. Lin-Liu et al. (2006)
  14. ^ Gregory, T.R. (2005). "Birds - Animal Genome Size Database". Genomesize.com. http://www.genomesize.com/statistics.php?stats=birds. Retrieved 2011-04-25. 
  15. ^ a b Madge et al. (2002)
  16. ^ Ray & Adams (2001)
  17. ^ Henninger (1906), OOS (2004)
  18. ^ NDGFD (1992)
  19. ^ "Roadkill: One from the road". London: The Independent. 2006-09-07. http://www.independent.co.uk/environment/roadkill-one-from-the-road-414972.html. Retrieved 2011-04-25. 
  20. ^ resourcesforhistory (2006)
  21. ^ a b h2g2 (2007)
  22. ^ "Pheasant History, Ecology & Biology". Pheasantsforever.org. http://www.pheasantsforever.org/page/1/pheasant.jsp. Retrieved 2012-03-11. 
  23. ^ Oregon Historical Society (20 August 2011). "Oregon pioneer Owen Nickerson Denny was about more than his birds". OregonLive.com. http://www.oregonlive.com/O/index.ssf/2011/08/oregon_pioneer_owen_nickerson.html. Retrieved 2012-03-11. 
  24. ^ Robertson (1997) p.124
  25. ^ BBC Four (2005)
  26. ^ "Game To Eat". Game To Eat. http://www.game-to-eat.co.uk/. Retrieved 2011-04-25. 
  27. ^ [Successful Farming, January (1988), p.g. 53]
  28. ^ E.g. Ohio: OOS (2004)
  29. ^ Robertson (1997) p.125
  30. ^ [1]

References

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Names and Taxonomy

Taxonomy

Comments: Sometimes considered as two separate species, P. COLCHICUS and P. VERSICOLOR. Within the COLCHICUS group, the Asiatic complex is sometimes treated as a species, P. TORQUATUS, distinct from the more western P. COLCHICUS (AOU 1983).

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