Overview
Distribution
Geographic Range
Ruffed grouse are found throughout Canada and in 38 of the 49 continental United States. In the east they extend as far south as northern Georgia. In the western states ruffed grouse are found along the western slopes of the Cascades into northern California and in the Rocky Mountains into Wyoming and central Utah. In the central United States isolated populations persist in the Dakotas and as far south as Arkansas. Populations have been introduced into Newfoundland and Nevada.
Biogeographic Regions: nearctic (Native )
- Ruffed Grouse Society, 2003. "Ruffed grouse facts" (On-line). Accessed February 14, 2005 at http://www.ruffedgrousesociety.org/ruffed_facts.asp.
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Global Range: RESIDENT: central Alaska and most of forested Canada south to northern California, central Idaho, central Utah, western South Dakota, Minnesota, central Arkansas, northern Georgia, and northeastern Virginia; introduced and established in Iowa, Newfoundland, Nevada, and part of Michigan (Godfrey 1966, AOU 1983, Crawford 1986). See Crawford (1986) for a map of distribution by subspecies. Was virtually extirpated from Indiana, Missouri, and Illinois, and populations were greatly reduced in Kentucky, Ohio, Tennessee, and parts of Montana, Saskatchewan, and Alberta. Reintroduced into Missouri and Indiana. Populations are limited in Illinois, Missouri, Nevada, and South Dakota.
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National Distribution
Canada
Origin: Native
Regularity: Regularly occurring
Currently: Present
Confidence: Confident
Type of Residency: Year-round
United States
Origin: Native
Regularity: Regularly occurring
Currently: Present
Confidence: Confident
Type of Residency: Year-round
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Physical Description
Morphology
Physical Description
The Ruffed Grouse is approximately 18 inches in length. The color is two-toned reddish-brown and spotted on the back, and yellowish with dark bars beneath. The tail has 18 broad feathers, which appear to be half-diamond shaped when spread. The tarsus is partly feathered.
Pearson, 1940.
Average mass: 644 g.
Average basal metabolic rate: 2.3894 W.
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Size
Ecology
Habitat
Habitat
The Ruffed Grouse prefers forested areas in rough, cold lands. It also prefers dim and quiet woods, deep thickets, or sheltered swamps. The Ruffed Grouse doesn't like open fields, and will rarely, if never, be found there.
Pearson,1940.
Terrestrial Biomes: forest
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Comments: Dense forest with some deciduous trees, in both wet and relatively dry situations from boreal forest (especially early seral stages dominated by aspen) and northern hardwood ecotone to eastern deciduous forest and oak-savanna woodland (AOU 1983). Young forest provides optimum conditions. See Crawford (1986) for additional information on habitat and cover requirements. Nests in forests or woodlands with some deciduous trees. Nest usually at base of tree, bush or stump. Drumming areas and broods usually in areas with high density of woody stems. Broods select habitats with abundant ground cover. In Wisconsin, females nested in areas in which they wintered following natal dispersal (Small and Rusch 1989).
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Migration
Non-Migrant: Yes. At least some populations of this species do not make significant seasonal migrations. Juvenile dispersal is not considered a migration.
Locally Migrant: No. No populations of this species make local extended movements (generally less than 200 km) at particular times of the year (e.g., to breeding or wintering grounds, to hibernation sites).
Locally Migrant: No. No populations of this species make annual migrations of over 200 km.
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Trophic Strategy
Food Habits
Over one-fourth of the Ruffed Grouse's diet is made up of fruit, such as thorn apples, blueberries, strawberries. It also eats wild and cultivated sunflower seeds, birch, poplar, and willow buds. The Christmas Fern is also a special food for the Ruffed Grouse.
Young Ruffed Grouse Chicks are primarily insectivorous, until they are old enough to care fo r themselves.
Birds of America, 1940.
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Comments: Young eat mainly insects and spiders. During summer insects may comprise about 30% of diet of adults. Adults also eat many herbaceous plants (especially in spring and summer), seeds, fruits (especially in fall and winter), nuts, flowers, buds, and leaves of trees and shrubs. In many areas, aspen (in boreal region, especially staminate buds and catkins), willow, and/or Rosaceae, or birch, alder, and hazelnut buds and catkins, are important food resources in winter and spring. See Crawford (1986) for further details.
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Associations
Known predators
Accipiter gentilis
Bubo virginianus
Homo sapiens
Based on studies in:
Canada: Manitoba (Forest)
This list may not be complete but is based on published studies.
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Known prey organisms
Corylus
Pyrola
Cornus
Aralia
Prunus
Amelanchier
Symphoricarpos
Pryola
Based on studies in:
Canada: Manitoba (Forest)
This list may not be complete but is based on published studies.
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General Ecology
Population densities may fluctuate dramatically. Some populations exhibit an approximately 10-year cycle of abundance. Spring density generally about 2-10/40ha; fall density up to 20-55/ha; densities highest in boreal forest region (Crawford 1986). Good habitats support densities better than 1 breeding male per 8 ha (Atwater and Schnell 1989).
Home range of brood about 6-19 ha, averages about 16 ha (40 acres). In Missouri, mean home range of adult males was 104 ha in fall-winter, 67 ha in spring-summer (Thompson and Fritzell 1989); Crawford 1986 reported examples of male and female home ranges that ranged from 2-12 ha. Mature and some immature males may defend a territory.
In Wisconsin, juvenile females in fall dispersed average of 4.8 km to winter range, males 2.1 km; in other areas dispersal distance often less than 2 km; juvenile males may wander extensively in spring (Small and Rusch 1989).
Usually roosts in small groups in winter. Fox predation often is an important cause of nesting failure. Other important predators include great horned owl and, in north, goshawk. In Alberta, predation by great horned owl declined when snowshoe hares were abundant (Rusch and Keith 1971). Shallow snow cover or icy crust on snow may reduce survival in winter by precluding access to subnivean shelter.
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Life History and Behavior
Life Expectancy
Lifespan/Longevity
Average lifespan
Status: wild: 102 months.
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Lifespan, longevity, and ageing
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Reproduction
Reproduction
The Ruffed Grouse is a ground nesting bird. The female lays one egg per day, until her clutch is complete. The average clutch is about 11 eggs. The female sits on the eggs, in the nest, until the eggs hatch, after 23 to 26 days. Fledging occurs after 8 to 10 weeks.
Pearson, 1940, USDA Forest Service 2001.
Average time to hatching: 24 days.
Average eggs per season: 11.
Average age at sexual or reproductive maturity (male)
Sex: male: 120 days.
Average age at sexual or reproductive maturity (female)
Sex: female: 120 days.
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Drumming and mating peak in mid-March in Missouri, early April in Indiana, and late April in Minnesota, early May in far north. The first eggs are laid in April or May, depending on the location (latitude). Clutch size is 4-19 (generally 9-12). Incubation, by female, lasts 23-24 days. Nestlings are precocial, downy; can fly in 10-12 days. Young are tended by female. Broods break up in fall when young about 84 days old; young disperse (Terres 1980), at about 120-125 days in Wisconsin (Small and Rusch 1989). In Alberta, about 50% of young survived from fall to spring (Rusch and Keith 1971). Single-brooded, but females may renest of first nesting attempt is unsuccessful. Sexually mature in one year; uncommonly lives more than 5 years. Cold wet weather in May/June may cause high losses among broods.
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Molecular Biology and Genetics
Molecular Biology
Barcode data: Bonasa umbellus
There are 7 barcode sequences available from BOLD and GenBank. Below is a sequence of the barcode region Cytochrome oxidase subunit 1 (COI or COX1) from a member of the species. See the BOLD taxonomy browser for more complete information about this specimen and other sequences.
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Download FASTA File
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Statistics of barcoding coverage: Bonasa umbellus
Public Records: 7
Species: 12
Species With Barcodes: 1
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Conservation
Conservation Status
IUCN Red List Assessment
Red List Category
Red List Criteria
Version
Year Assessed
Assessor/s
Reviewer/s
Contributor/s
Justification
History
- 2008Least Concern
- 2004Least Concern
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Conservation Status
The Ruffed Grouse is hunted for sport, but it is far from being endangered. The only real requirement that the Ruffed Grouse needs is a forested region, so this is another animal that could be affected by extensive deforestation.
US Migratory Bird Act: no special status
US Federal List: no special status
CITES: no special status
State of Michigan List: no special status
IUCN Red List of Threatened Species: least concern
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National NatureServe Conservation Status
Canada
Rounded National Status Rank: N5 - Secure
United States
Rounded National Status Rank: N5 - Secure
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Management
Preserve Selection and Design Considerations: A management unit for providing total food and cover needs should be at least 10-16 ha (25-40 acres).
Management Requirements: Healthy populations may be able to withstand a 50% harvest without affecting spring density (Crawford 1986). See Williamson (no date), Crawford (1986), and Atwater and Schnell (1989) for information on habitat management (e.g., cutting, plantings, burning, grazing, etc.).
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Relevance to Humans and Ecosystems
Benefits
Economic Importance for Humans: Negative
Some farmers are troubled by these birds, because they are primarily fruit eaters.
Leonard, 1973.
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Economic Importance for Humans: Positive
The Ruffed Grouse is hunted for sport, so areas that have very dense populations of the bird benefit from extra tourism during the hunting season
Also, The Ruffed Grouse hatchlings are mainly insectivorous, so the insect populations in some area decrease shortly after the hatching season.
Peterson, 1980.
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Economic Uses
Comments: The third most heavily harvested galliform in North America; 70% of harvest occurs in Minnesota, Michigan, New York, Pennsylvania, and Wisconsin (Johnsgard 1973).
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Wikipedia
Ruffed Grouse
The Ruffed Grouse (Bonasa umbellus) is a medium-sized grouse occurring in forests from the Appalachian Mountains across Canada to Alaska. It is non-migratory.
The Ruffed Grouse is frequently referred to as a "partridge". This is technically wrong—partridges are unrelated phasianids, and in hunting may lead to confusion with the Grey Partridge, it is a bird of woodlands, not open areas.[1]
The Ruffed Grouse is also the state bird of Pennsylvania.
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Description
These chunky, medium-sized birds weigh from 450–750 g (0.99–1.7 lb), measure from 40 to 50 cm (16 to 20 in) in length and span 50–64 cm (20–25 in) across their short, strong wings.[2] Ruffed Grouse have two distinct morphs, grey and brown. In the grey morph, the head, neck and back are grey-brown; the breast is light with barring. There is much white on the underside and flanks, and overall the birds have a variegated appearance; the throat is often distinctly lighter. The tail is essentially the same brownish grey, with regular barring and a broad black band near the end ("subterminal"). Brown-morph birds have tails of the same color and pattern, but the rest of the plumage is much more brown, giving the appearance of a more uniform bird with less light plumage below and a conspicuously grey tail. There are all sorts of intergrades between the most typical morphs; warmer and more humid conditions favor browner birds in general.
The ruffs are on the sides of the neck in both sexes. They also have a crest on top of their head, which sometimes lies flat. Both sexes are similarly marked and sized, making them difficult to tell apart, even in hand. The female often has a broken subterminal tail band, while males tend to have unbroken tail bands, though the opposite of either can occur. Females may also do a display similar to the male. Another fairly accurate sign is that rump feathers with a single white dot indicate a female; rump feathers with more than one white dot indicate a male.
Ecology
Like most grouse, they spend most of their time on the ground, and when surprised, may explode into flight, beating their wings very loudly. Mixed woodland rich in aspen seems to be particularly well-liked. These birds forage on the ground or in trees. They are omnivores, eating buds, leaves, berries, seeds, and insects. According to Johnson:
"More than any other characteristic, it is the ruffed grouse's ability to thrive on a wide range of foods that has allowed it to adapt to such a wide and varied range of habitat on this continent. A complete menu of grouse fare might itself fill a book. One grouse crop yielded a live salamander in a salad of watercress. Another contained a small snake."[3]
Hunting
"Hunting of the ruffed grouse is common in the northern and far western United States as well as Canada, Often done with shotguns. Dogs may also be used. Hunting of the ruffed grouse is usually very rigorous. This is because the grouse spends most of its time in thick brush, aspen stands, and second growth pines. It is also very hard to detect a foraging grouse bobbing about in the thicket. This is because their deceptive plumage hides them very well in a variety of conditions, though when snow is present it may become a lot easier. Like other forest creatures, the ruffed grouse will maintain trails through the underbrush and pines. These can often be found by looking for the feathers of the bird on the ground and on twigs at the edges of its trail. Hunting of the ruffed grouse requires a good ear and lots of stamina as you will be constantly walking and listening for them in the leaves." - Joseph B. Barney
Ruffed Grouse will frequently seek gravel and clover along road beds during early morning and late afteroon. These areas are good areas to walk during this time to flush birds. Also, grouse use sandy road beds to dust their feathers to rid themselves of skin pests. Dusting sites are visible as areas of disturbed soils with some signs of feathers. Birds will return to these spots during the late afternoon to dickbathe in dust and also socialize and mate. W. Macht
The Ruffed Grouse population has a cycle, and follows the cycle no matter how much or how little hunting there is. The cycle has puzzled scientists for years, and is simply referred to as the "grouse cycle."
Minnesota is the top Ruffed Grouse–producing state in the U.S.[4]
See also
Footnotes
- ^ Ruffed Grouse, Animal Diversity
- ^ [1]
- ^ Johnson (1995): p.37
- ^ http://www.dnr.state.mn.us/hunting/grouse/index.html
References
| Wikimedia Commons has media related to: Bonasa umbellus |
- BirdLife International (2004). Bonasa umbellus. 2006. IUCN Red List of Threatened Species. IUCN 2006. www.iucnredlist.org. Retrieved on 11 May 2006.
- Henninger, W. F. (1906): A preliminary list of the birds of Seneca County, Ohio. Wilson Bulletin 18(2): 47-60. DjVu fulltext PDF fulltext
- Johnson, Don L. (1995): Grouse & Woodcock: A Gunner's Guide. Krause Publications. ISBN 0-87341-346-6.
- Ohio Ornithological Society (OOS) (2004): Annotated Ohio state checklist. Version of April 2004. PDF fulltext
- State Symbols of Pennsylvania: State Bird, The Ruffed Grouse PDF fulltext
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Names and Taxonomy
Taxonomy
Comments: Ellsworth et al. (1995) examined phylogenetic relationships among North American grouse based on mtDNA data and found that Falcipennis canadensis is more closely related to Bonasa umbellus (ruffed grouse) than to Dendragapus obscurus (blue grouse), which is allied with Lagopus (ptarmigan) and Tetrao (capercaillie, a European grouse).
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