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Overview

Brief Summary

Biology

Steller's sea eagles start to build their large, bulky nests in the trees in late February and early March (3). The first eggs are laid in mid-April, and clutch size varies from 1 to 3 eggs; hatchlings emerge in mid-May to mid-June and begin to fly by August and early September (3).  These large birds feed predominately on salmon (Onchorhynchus spp.), which are taken both dead and alive. Prey is usually caught by swooping from perches located at the waters' edge, or from circling and diving down; occasionally birds will stand in the shallows to catch fish (5). The large, powerful bill is perfectly adapted to ripping and tearing at flesh and these birds will also prey on other fish and the carcasses of animals such as seals and sea lions (3). Where there are large congregations of prey such as salmon, groups of eagles will gather and individuals will often attempt to steal food from each other in a behaviour known as 'kleptoparasitism' (3).
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Description

Steller's sea eagle is one of the largest of the sea and fish eagles of the genus Haliaeetus (3). These large blackish-brown birds have an enormous, strongly arched yellow bill (3). The feathers on the shoulders, tail and legs are white (2), and females are generally the larger sex (3).
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Distribution

Steller's sea eagles are native to eastern Russia, specifically, the Sea of Okhotsk and the Kamchatka Peninsula. They are frequent winter migrants south to the Japanese Islands of Kuril and Hokkaido and have been seen as far south as eastern China and Korea. Vagrant individuals have also been spotted in Taiwan and the United States.

Biogeographic Regions: palearctic (Native )

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Range Description

Haliaeetus pelagicus breeds on the Kamchatka peninsula, the coastal area around the Sea of Okhotsk, the lower reaches of the Amur river (south to the Gorin river) and on northern Sakhalin and Shantar, Russia. A few hundred winter in Kamchatka, the northern Sea of Japan, and the coast of Okhotsk, but most (c.2,000) winter in the southern Kuril islands and Hokkaido, Japan. It is an uncommon winter visitor to north-eastern China, North Korea and South Korea. Declining breeding success has been noted in the inland river populations of Magadan district, Russia, from 1991 to 2009, with a slow increase in the breeding success of coastal populations over the same period, suggesting that they can be considered sink and source populations respectively (Potapov et al. 2010). Its total population is estimated at c.5,000 mature individuals and declining overall.

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Range

Coastal e Siberia; winters to China, Korea, Japan and Ryukyu Is..
  • Clements, J. F., T. S. Schulenberg, M. J. Iliff, D. Roberson, T. A. Fredericks, B. L. Sullivan, and C. L. Wood. 2014. The eBird/Clements checklist of birds of the world: Version 6.9. Downloaded from http://www.birds.cornell.edu/clementschecklist/download/

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Range

Steller's sea eagles breed in eastern Russia, around the Sea of Okhotsk and on the Kamchatka Peninsula. A small number of birds remain in Kamchatka over the winter but the majority fly south to the Japanese Islands of Kuril and Hokkaido (2). This species is also occasionally seen in China and North and South Korea (2).
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Physical Description

Morphology

Steller's sea eagles are large eagles with dark brown to black feathers on the majority of the body and white on the shoulders, thighs, and crown. They have wedged-shaped, white tails, very large yellow beaks, and sharp, yellow talons. Average mass is 6 kilograms in males and 9 kilograms in females. Body length of both males and females ranges between 85 and 94 centimeters with average wingspans of females around 136 centimeters and males around 118 centimeters.

Average mass: males - 6, females - 9 kg.

Range length: 85 to 94 cm.

Average wingspan: males - 118, females - 136 cm.

Other Physical Features: endothermic ; homoiothermic; bilateral symmetry

Sexual Dimorphism: female larger

  • Brown, L., D. Amadon. 1989. Eagles, Hawks and Falcons of the World. Edison, New Jersey: The Wellfleet Press.
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Ecology

Habitat

Steller's sea eagles breed along sea coasts or near large rivers with mature trees. Sightings very far inland are rare, as they prefer sea coasts that are dotted with estuaries and river mouths. They nest on large, rocky outcroppings or at the tops of large trees. Steller's sea eagles are generally found at elevations ranging from sea level to approximately 100 m.

Migrating Steller's sea eagles winter along rivers in Japan and occasionally move to mountainous inland areas as opposed to the sea coast. They are also occasionally seen over and perching on sea ice in northern waters.

Range elevation: 0 to 100 m.

Habitat Regions: temperate ; terrestrial ; saltwater or marine ; freshwater

Aquatic Biomes: rivers and streams; coastal

Wetlands: marsh ; swamp

Other Habitat Features: riparian ; estuarine

  • Collar, N. 2001. Threatened Birds of Asia: The BirdLife International Red Data Book. Cambridge, UK: Birdlife International.
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Habitat and Ecology

Habitat and Ecology
It breeds on sea coasts and inland near larger rivers (mostly on lower stretches) or lakes, where there are stands of mature trees. In the Magadan district of Russia, successful breeding pairs along coasts appear to produce more fledged chicks than successful pairs on rivers, and average brood size is larger for coastal pairs (Potapov et al. 2010). During the autumn birds forage along rivers where dead salmon are abundant. During mid-winter, birds in Russia tend to remain on the coast, except some that winter in Kamchatka along inland rivers fed by hot springs and at Lake Kurilskoye (M. McGrady et al. in litt. 2012), while those wintering in Japan mainly stay near freshwater, but c.35% move to mountainous areas where many feed on deer carcasses (Ueta et al. 2003).


Systems
  • Terrestrial
  • Freshwater
  • Marine
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Inhabits coastal cliffs and estuaries; further inland, these birds are associated with river and lakeshore forests (5).
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Trophic Strategy

The main prey of Steller's sea eagles are salmon, taken either dead or alive. When salmon is scarce or not available, other food resources are taken, ranging from invertebrates like crabs and mussels to gulls, small mammals, and carrion. Three types of hunting behaviors have been observed, hunting from a perch, hunting on the wing while circling 6 to 7 meters above the water, and hunting in shallow water. Kleptoparasitism has also been observed when feeding occurs in groups and food is abundant, adults benefit the most from this behavior.

Animal Foods: birds; mammals; fish; carrion ; mollusks; aquatic crustaceans

Primary Diet: carnivore (Piscivore , Scavenger )

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Associations

Steller's sea eagles are important predators of salmon and other prey in their native ecosystems.

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There are no known predators of adult Steller's sea eagles. Eggs and hatchlings are commonly preyed on by arboreal mammals that gain access to nests, such as martens, and by crows.

Known Predators:

  • crows (Corvus)
  • martens (Martes)

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Life History and Behavior

Behavior

Steller's sea eagles communicate mainly through various vocalizations. A deep, barking cry is commonly heard. During mating displays a loud, gull-like call is used.

Communication Channels: acoustic

Perception Channels: visual ; tactile ; acoustic ; chemical

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Life Expectancy

Little is known about the lifespan of Steller's sea eagles, but it is thought to be similar to that of their close relatives, white-tailed sea eagles (Haliaeetus albicilla), which live 20 to 25 years in the wild.

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Reproduction

Steller's sea eagles are monogamous, they are often seen in breeding pairs throughout the breeding season, usually lasting from February through August. Both males and females secure their own breeding territories early in the season and nest building occurs in February or March. Displaying begins in March and consists of soaring high above the breeding area while calling.

Mating System: monogamous

Both male and female Steller's sea eagles reach sexual maturity by six or seven years. Breeding occurs seasonally between February and August, beginning with nest building in February and March. Typically, a pair will maintain two to four nests in one breeding territory and use alternate nests from year to year. Nests are most often built on rocky cliffs or in large trees out of thick branches and can reach a size of two meters across and two to four meters thick. The average clutch size is 2 but ranges from 1 to 3. The egg-laying period normally lasts from April through May, and the typical incubation period is 38 days. Eggs hatch between May and June, with fledging taking around 70 days. Chicks leave nests by August or September.

Breeding interval: Steller's sea eagles breed once yearly.

Breeding season: Breeding occurs from February through August.

Range eggs per season: 1 to 3.

Range time to hatching: 38 to 45 days.

Average fledging age: 70 days.

Average time to independence: 70 days.

Range age at sexual or reproductive maturity (female): 6 to 7 years.

Range age at sexual or reproductive maturity (male): 6 to 7 years.

Key Reproductive Features: iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous

At this time, little is known about the parental investment of Steller's sea eagles. Both parents contribute to raising offspring to independence.

Parental Investment: altricial ; pre-fertilization (Provisioning, Protecting: Female); pre-hatching/birth (Provisioning: Female, Protecting: Male, Female); pre-weaning/fledging (Provisioning: Male, Female, Protecting: Male, Female); pre-independence (Provisioning: Male, Female, Protecting: Male, Female)

  • Collar, N. 2001. Threatened Birds of Asia: The BirdLife International Red Data Book. Cambridge, UK: Birdlife International.
  • 2007. "BirdLife International Species factsheet: Haliaeetus pelagicus" (On-line). Accessed March 17, 2008 at http://www.birdlife.org.
  • 1999. "Species Synopsis Stellar Sea Eagle" (On-line). Accessed March 24, 2008 at http://www.fadr.msu.ru/o-washinet/spsynop.html.
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Molecular Biology and Genetics

Molecular Biology

Barcode data: Haliaeetus pelagicus

The following is a representative barcode sequence, the centroid of all available sequences for this species.


There is 1 barcode sequence available from BOLD and GenBank.

Below is the sequence of the barcode region Cytochrome oxidase subunit 1 (COI or COX1) from a member of the species.

See the BOLD taxonomy browser for more complete information about this specimen.

Other sequences that do not yet meet barcode criteria may also be available.

ATCTTCGGCGCCTGAGCTGGCATAGTCGGCACCGCCCTC---AGTTTACTCATTCGCGCAGAACTCGGCCAACCAGGCACACTCCTAGGCGAC---GACCAAATCTATAACGTAATCGTCACCGCACATGCCTTCGTAATAATCTTCTTCATAGTTATACCCATCATAATCGGAGGATTCGGAAACTGACTCGTCCCGCTCATA---ATTGGCGCCCCCGACATAGCCTTCCCACGCATAAACAACATAAGCTTCTGACTACTTCCTCCCTCCTTCCTCCTCTTACTAGCCTCCTCAACTGTAGAAGCAGGAGCTGGCACCGGATGAACTGTTTATCCCCCATTAGCAGGCAACATGGCCCATGCCGGAGCCTCAGTAGACTTA---GCCATCTTCTCCTTACACCTAGCTGGAATCTCATCCATTCTAGGGGCAATCAACTTTATCACAACCGCTATCAACATAAAACCCCCAGCCCTCTCCCAATACCAAACACCCTTATTTGTATGATCCGTCCTTATCACCGCCGTCCTACTACTACTCTCTCTCCCAGTCCTAGCCGCT---GGCATCACTATACTACTTACAGATCGAAACCTCAACACAACATTCTTCGACCCTGCTGGTGGAGGTGACCCCATCCTATACCAGCATCTCTTCTGATTCTTCGGACACCCGGAAGTCTACATCCTAATTCTACC
-- end --

Download FASTA File

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Statistics of barcoding coverage: Haliaeetus pelagicus

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 2
Specimens with Barcodes: 7
Species With Barcodes: 1
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Conservation

Conservation Status

Steller's sea eagles have a limited population size of around 5,000 individuals. Populations are in steady decline. Potential causes of declining populations are habitat degradation due to an increase in industry and logging, overfishing of key prey items, and pollution.

Steller's sea eagles are listed as vulnerable on the IUCN Red List of Threatened Species because of their small and declining population sizes. Steller's sea eagles are legally protected in Russia, Japan, China, and South Korea with key habitat areas being established as nature reserves throughout Russia and Japan. They are also protected under Appendix II of the Convention on International Trade of Endangered Species.

US Migratory Bird Act: protected

US Federal List: no special status

CITES: appendix ii

State of Michigan List: no special status

IUCN Red List of Threatened Species: vulnerable

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IUCN Red List Assessment


Red List Category
VU
Vulnerable

Red List Criteria
C2a(ii)

Version
3.1

Year Assessed
2013

Assessor/s
BirdLife International

Reviewer/s
Butchart, S.

Contributor/s

Justification
This species has a small, regionally declining population as a result of habitat degradation, pollution, poisoning by lead shot, and over-fishing. It therefore qualifies as Vulnerable.


History
  • 2012
    Vulnerable
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Status

Classified as Vulnerable (VU - C1) on the IUCN Red List 2002 (1). Listed on Appendix II of CITES (4), and Appendix I of the Convention on Migratory Species (CMS or Bonn Convention) (6).
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Population

Population
The global population is estimated to number c.4,600-5,100 individuals, including c.1,830-1,900 breeding pairs (M. McGrady et al. in litt. 2012), assumed to be equivalent to c.3,600-3,800 mature individuals.

Population Trend
Decreasing
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Threats

Major Threats
In Russia, it is threatened by habitat alteration during the development of hydroelectric power projects, proposed large-scale coastal and offshore developments for the petrochemical industry, and logging for timber. Industrial pollution of rivers and high levels of DDT/DDE, PCBs and heavy metals are further threats. Over-fishing has caused a decline of fish stocks in Russia and Japan which has led to an increasing tendency of birds on Hokkaido to move inland and scavenge on sika deer carcasses left by hunters, exposing them to a risk of lead poisoning through ingestion of lead shot.

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Steller's sea eagles are under threat from a variety of factors. In Russia, the most pertinent threat is habitat loss caused by hydro-electro schemes, together with logging for timber and coastal development (5). Salmon stocks have been depleted in much of the range of this eagle, which in turn threatens population numbers (2).
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Management

Conservation Actions

Conservation Actions
Conservation Actions Underway
CITES Appendix II, CMS Appendix I and II. It is legally protected in Russia, Japan, China and South Korea. It is monitored in several protected areas in Russia, including the Magadan State Nature Reserve, Kronotski State Reserve, Lake Krontskoea Wildlife Refuge and Kava Wildlife Refuge (Magadan), the Orel' and Udyl' Wildlife Refuges and Dzhugdzhurskiy, Shantarsky and Komsomol'ski Nature Reserves (Khabarovsk), the Poronayskiy Nature Reserve (Sakhalin), and the Kuril'ski Nature Reserve (Kuril Islands). In Japan, the key wintering grounds on Hokkaido, Shiretoko and Furen-ko are designated as National Wildlife Protection Areas.

Conservation Actions Proposed
Conduct surveys to obtain an up-to-date population estimate. Monitor population trends through regular surveys. Minimise the damaging effects of industrial development in its Russian breeding grounds. In some wintering areas, consider establishing special artificial feeding-sites. Ensure regular sampling of the environment and the species for DDT/DDE, PCBs and other pollutants in Khabarovsk and Magadan, and for lead in Japan. Protect important salmon spawning grounds. Encourage sustainable management of key fish stocks. Preserve potential nest trees in river valleys within 30 km of the sea (M. McGrady et al. in litt. 2012). Establish no disturbance zones in estuaries where conflicts exist between fishers and eagles (M. McGrady et al. in litt. 2012). Develop a captive breeding programme to support future reintroduction and supplementation efforts.

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Conservation

The magnificent Steller's sea eagle is protected within the countries of its range and internationally by its listing on the Convention on Migratory Species (CMS) (2). In addition, international trade is restricted by the placing of this species on Appendix II of the Convention on International Trade in Endangered Species (CITES) (4). In Russia, these sea eagles are found in a number of reserves where their populations are monitored; the important wintering grounds on the islands of Hokkaido, Shiretoko and Furen-ko in Japan are National Wildlife Protection Areas (2).
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Relevance to Humans and Ecosystems

Benefits

Steller's sea eagles have been known to remove mammals from commercial traps set by humans during harsh winters, causing some harm to that industry.

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There are no described benefits of Steller's sea eagles to humans.

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Wikipedia

Steller's sea eagle

The Steller's sea eagle (Haliaeetus pelagicus) [2] is a large bird of prey in the family Accipitridae. It is an eagle that lives in coastal northeastern Asia and mainly preys on fish and water birds. On average, it is the heaviest eagle in the world, at about 5 to 9 kilograms (11 to 20 lb), but may lag behind the harpy eagle (Harpia harpyja) and Philippine eagle (Pithecophaga jefferyi) in some standard measurements.[3] This bird is named after the German naturalist Georg Wilhelm Steller.[4]

Etymology[edit]

This species was first described as Aquila pelagica by Peter Simon Pallas, in either 1811 or 1826 depending on the source. Subsequently, many generic and specific names have been variously spelled, e.g., Haliaetus pelagicus, Haliaetos pelagica, Faico leucopterus, Faico imperator, Thalassaetus pelagicus, Thalassaetus macrurus, Haliaeetus macrurus and most recently Thallasoaetus pelagicus. Besides its normal common name, the species has sometimes been referred to as the Pacific eagle or White-shouldered eagle. In Russian, the eagle has been called morskoi orel (sea eagle), pestryi morskoi orel (mottled sea eagle) or beloplechii orlan (white-shouldered eagle). In Japanese, it is called 0-washi (large eagle or great eagle).[5]

Description[edit]

A Steller's sea eagle flying with a fish in Hokkaido, Japan
A Steller's sea eagle near Rausu, Hokkaido, Japan
Detail of head – taken at the Cincinnati Zoo
A falconer holding a Steller's sea eagle in the Yorkshire Dales, England

Size[edit]

Steller's Sea-eagle is the biggest bird in the genus Haliaeetus and is one of the largest raptors overall. Females may vary in weigh from 6,195 to 9,500 g (13.658 to 20.944 lb) while males being rather lighter with a weight range from 4,900 to 6,800 g (10.8 to 15.0 lb).[3][6][7] The average weight is variable, possibly to due to seasonal variation in food access or general condition of eagles, but has been reported from as high as a mean mass of 7,757 g (17.101 lb) to a median estimate weight of 6,250 g (13.78 lb), excluding expired eagles who were poisoned by lead and endured precipitous weight loss by the occasion of their deaths.[8][3][6][7][9] At its average weight, the Steller's seems to outweigh the average Harpy by approximately 500 g (1.1 lb) and the average Philippine eagles by more than 1,000 g (2.2 lb).[3][7][10] The Steller's sea eagle can range in total length from 85 to 105 cm (2 ft 9 in to 3 ft 5 in), apparently males average about 89 cm (2 ft 11 in) in length, while females average about 100 cm (3 ft 3 in), marginally shorter (a few mm) on average than the harpy eagle and about 65 mm (2.6 in) shorter than the Philippine eagle.[3][8] The wingspan is from 1.95 to 2.5 m (6 ft 5 in to 8 ft 2 in) and the wing chord measurement is 560 to 680 mm (22 to 27 in).[3][5][11] The Steller's sea eagle's wingspan is one of the largest of any living eagle, at an median of 2.13 m (7 ft 0 in) per Ferugson & Lees (2001) or a median of 2.2 m (7 ft 3 in) per Saito (2009).[3][8] Its closest rivals are the closely related white-tailed eagle (Haliaeetus albicilla), at reported median wingspans of 2.1 and 2.18 m (6 ft 11 in and 7 ft 2 in) and the unrelated wedge-tailed eagle (Aquila audax), at reported average wingspans of 2.04 and 2.23 m (6 ft 8 in and 7 ft 4 in), nonetheless both other eagles are rather smaller in weight and total length.[3][8][12]

Standard Measurements and Physiology[edit]

As in most Haliaeetus eagles, the tarsus and tail are relatively short compared to other very large eagles at 95–100 mm (3.7–3.9 in) and 320–390 mm (13–15 in) in length, respectively, the Philippine eagle besting it by up to 40 mm (1.6 in) and 110 mm (4.3 in) apparently.[3][13] In all sea and fish eagle, the toes are relatively short and stout, with the bottom of the foot covered in spiracles and the talons being relatively shorter and more strongly curved than in comparably sized eagles found in forests and fields, such as the "booted eagle" group (i.e. Aquila) or "harpy eagles", all of these specializations developed in the aid of capturing fish rather than medium-sized mammals and large birds, although clearly these are not excluded from capture.[3][13] Perhaps the most noted physical feature of the Steller's sea eagle, other than its overall great size, is its extremely large bill and prominent head. The skull is around 14.6 cm (5.7 in) in total length, the culmen is from 62 to 75 mm (2.4 to 3.0 in) and the bill from the gape to the tip is around 117 mm (4.6 in).[14][15] The Steller's sea eagle's bill is probably the largest of any living eagle, just surpassing to the Philippine eagle with a sole known culmen measurement (from a mature female) of 72.2 mm (2.84 in), and are similar in robustness (if slightly shorter in culmen length) to those of the largest accipitrids, the Old World vultures.[3][14][16]

Coloring[edit]

The mature Steller's sea eagle is dark brown to black over the majority of its body, with strongly contrasting white on the lesser and median upper-wing coverts, underwing coverts, thighs, under-tail coverts and tail. They have wedge-shaped, white tails that are relatively longer than those of the white-tailed eagle. The bold, pied coloration of adults may play some part in social hierarchies with other eagles of their own species during the non-breeding season, although this has not been extensively studied.[3][13] The eyes, the bill and the feet of adults are all yellow in colouration. Two subspecies have been named: The relatively widespread nominate pelagicus and the virtually unknown H. p. niger.[17] The latter name was given to the population which lacked white feathers except for the tail and supposedly was resident all year in Korea. Last seen in 1968 and long believed to be extinct, a female matching H. p. niger in appearance was born in captivity in 2001. Both its parent were "normal" in appearance, indicating that H. p. niger is an extremely rare morph rather than a valid subspecies, as had already been suggested earlier.[18][19]

The first down plumage of new nestlings is silky white, though they soon turn a smoky brown-grey. As in other sea eagles, remiges and retrices of the first-year plumage are longer than adults. Juvenile plumage is largely a uniform dark brown with occasional grey-brown streaking about the head and the neck, white feather bases and light mottling on the retrices. The tail of the immature eagle is white with black mottling basally.[13][9] The young Steller's sea eagle has a dark brown iris, whitish legs and blackish-brown beak. Through at least three intermediate plumages, mottling in the tail decreases, body and wing feathering acquires a bronze cast, and the eye and bill lighten in colour. Definitive plumage is probably reached in the fifth year of life, based on fragmentary data from captives. First and intermediate plumages are difficult to distinguish from those of the white-tailed eagle, which occurs in the entire breeding range of the Steller's.[5]

Voice[edit]

Steller's sea eagles are not extensively known for their voices but are known to make a deep barking cry, ra-ra-ra-raurau, in aggressive interactions. Their call is similar to the white-tailed eagles but deeper.[13] During the display at the beginning of the breeding season, they have been heard to make calls to each that sound like very loud, deep-voiced gulls.[13][20]

Systematics and taxonomic status[edit]

The relationships of Steller's Sea-eagle are not completely resolved. mtDNA cytochrome b sequence data tentatively suggests that this species's ancestors diverged early in the colonization of the Holarctic by sea eagles. This is strongly supported by morphological traits such as the yellow eyes, beak, and talons shared by this species and the other northern sea-eagles, the White-tailed and bald eagles (Haliaeetus leucocephalus), and biogeography.[21] It is unique among all sea eagles in having a yellow bill even in juvenile birds, and possessing 14, not 12, rectrices.

Distribution and habitat[edit]

Steller's sea eagle breeds on the Kamchatka Peninsula, the coastal area around the Sea of Okhotsk, the lower reaches of the Amur River and on northern Sakhalin and the Shantar Islands, Russia. The majority of birds winter farther south, in the southern Kuril Islands, Russia and Hokkaidō, Japan. That being said, the Steller's Sea-eagle is less vagrant than the white-tailed eagle, usually lacking the long-range dispersal common in juveniles of that species.[22][23] Vagrant eagles have been found in North America, at locations including the Pribilof Islands and Kodiak Island, inland to as far as Peking in China and Yakutsk in Russia's Sakha Republic and south to as far as Taiwan but these are considered to be individual eagles that have strayed far from the species' typical range.[13][24]

The large body size (see also Bergmann's rule) and distribution of the Steller's sea eagle suggests that it is a glacial relict, meaning that it evolved in a narrow subarctic zone of the northeasternmost Asian coasts, which shifted its latitude according to ice age cycles, and never occurred anywhere else. This bird nests in two habitats: along sea coasts and alongside large rivers with mature trees. They nest on large, rocky outcroppings or at the tops of large trees. Usually areas with large Erman's birches (Betula ermanii) and floodplain forests of larches, alders, willows and poplar seem to be the preferred nesting spots. Some eagles, especially those that nest in sea coast, may not migrate. The timing, duration and extent of migration depends on ice conditions and food availability. On Kamchatka, eagles overwinter in forests and river valleys near the coast, but are irregularly distributed over the peninsula. Most wintering birds there appear to be residential adults. Steller's sea eagles that do migrate fly down to winter in rivers and wetlands in Japan but will occasionally move to mountainous inland areas as opposed to the sea coast. Each winter, drifting ice on the Sea of Okhotsk drives thousands of eagles south. Ice reaches Hokkaido in late January. Eagle numbers peak in the Nemuro Strait in late February. On Hokkaido, eagles concentrate in coastal areas and on lakes near the coast, along with substantial numbers of white-tailed eagles. Eagles depart between late March and late April, adults typically leaving before immatures. Migrants tend to follow sea coasts and are usually observed flying singly. In groups, migrants are typically observed flying 100–200 m (330–660 ft) apart. On Kamchatka, most migrants are birds in transitional plumages. They are also occasionally seen flying over the Northern ocean or perching on sea ice during the winter.[5][24]

Diet[edit]

An injured Pacific cod, an example of one of the many fish that these eagles subsist on
A Slaty-backed Gull, one of the primary avian species hunted by Steller's Sea Eagles

The Steller's sea eagle mainly feeds on fish. Their favored prey in river habitats are salmon (Oncorhynchus spp.) and trout.[25] Among these, pink salmon (O. gorbuscha) and chum salmon (O. keta) are reportedly favored, sometimes intensely supplemented by grayling (Thymallus thymallus) and three-spined stickleback (Gastrossteus aculeatus).[26] While pink and chum salmon average approximately 2,200 and 5,000 g (4.9 and 11.0 lb) in mature mass, respectively, the Steller's sea eagle not infrequently predates fish of up to 6,000 to 7,000 g (13 to 15 lb).[14] In coastal areas, nesting eagles may feed on Bering wolffish (Anarchichas orientalis), Hemitripterus villosus, Aptocyclus ventricosus and Myoxocephalus spp.[26] Like most Haliaeetus eagles, they hunt fish almost exclusively in shallow water. Relatively large numbers of these normally solitary birds can be seen congregating on particularly productive spawning rivers in August through September due to an abundant food supply.[24] On Kamchatka, aggregations of as many as 700 eagles have been reported, though much smaller groups are the norm.[5] In summer, live fish, typically in the range of 20 to 30 cm (7.9 to 11.8 in) in length, are fed to the young at the nest. Normally the parents will catch about 2 or 3 fish for the young to eat each day. In autumn, when many salmon die after spawning, dead fish tend to be consumed more often than live ones, and these are the main food for Steller's sea eagles who overwinter in inland rivers with non-frozen waters.[13] On Hokkaido, eagles are attracted by abundant Pacific cod (Gadus macrocephalus) which peak in the Rausu Sea and the Nemuro Straits in February. This resource supports an important commercial fishery which in turn helps to support eagles.[5] Alaska pollock (Theragra chalcogramma), along with the cod, is the most important food source for wintering eagles in Japan.[3]

Fish comprise about 80% of the diet of eagles nesting in the Amur River. Elsewhere, other prey can comprise almost an equally important portion of the diet.[5] Non-piscivore prey consists largely of water-dwelling birds, including ducks, geese, swans, cranes, herons and gulls. Along the sea coast and in Kamchatka, water birds are the most common prey for Steller's sea eagles.[27] Among bird prey, this eagle has shown a strong local preference for slaty-backed gulls (Larus schistisagus).[27] Common and thick-billed murres (Uria aalge & U. lomvia respectively) dominated the diet around the Sea of Okhotsk, followed by black-legged kittiwakes (Rissa tridactyla), slaty-backed gulls, crested auklets (Aethia cristatella) and pelagic cormorant (Phalacrocorax capillatus).[26] Small chicks of murres and cormorants were sometimes taken alive in Russia and brought back to nests and independently feed on remains of fish in the eagle's nests until they were killed themselves.[26] In Russia, upland bird species, black-billed capercaillie (Tetrao parvirostris) and Willow and rock ptarmigan (Lagopus lagopus & L. muta) can be an important prey species. Grouse are not typically taken by other Haliaeetus species.[13][28] Other landbirds hunted by Steller's sea eagles have included short-eared owls (Asio flammeus), snowy owl (Bubo scandiaca), carrion crow (Corvus corone) and common raven (Corvus corax), as well as (rarely) smaller passerines.[26] In one case, a Steller's sea eagle was observed feeding on a rare prey item in the Northern Hemisphere, a great albatross (Diomedea), as that genus nests in the sub-Antarctic oceans.[3] This sea eagle may supplement its diet with various mammals (especially hares[3][13]), crabs, mussels, Nereis worms and squid when given the opportunity.[26][29] Mammalian carnivores are apparently readily hunted. These include sable (Martes zibellina), American mink (Neovison vison), Arctic fox (Vulpes lagopus), red fox (Vulpes vulpes) and small domestic dogs (Canis lupus domesticus).[13][26] Smaller mammalian more rarely recorded prey includes northern red-backed vole (Clethrionomys rutilus) and tundra vole (Microtus oeconomus).[26] Carrion, especially that of mammals, is readily eaten during the winter. Around 35% of eagles wintering in Japan move inland and feed largely on mammalian carcasses, predominantly sika deer (Cervus nippon).[30] In winter, immature Steller's sea eagles may frequent slaughterhouses to pirate bits of offal.[13] This eagle has been recorded preying occasionally on young seals. It was estimated in one study (Brown & Amadon), that some seal pups carried off in flight by the eagles weighed at least 9.1 kg (20 lbs), which (if true) would be the greatest load carrying ever known for a bird; however the prey weights were not verified.[31] Often seals and sea lion of any size are eaten as carrion and, using the huge bill, may be dismembered where found rather than flown with.[32]

Most often, Steller's sea eagles hunt from a perch in a tree or rocky ledge located 5–30 m (16–98 ft) above the water. When prey is spotted, the bird dives from its perch. Eagles may also hunt on the wing, while circling 6–7 m (20–23 ft) above the water. Again, prey is captured by diving. Eagles sometimes hunt by standing in or near shallow water on a sandbank, spit, or ice-flow, grabbing passing fish. It is reported that, compared to its white-tailed and bald eagle cousins, the Steller's sea eagle is a more "aggressive, powerful and active" raptor.[13] Where feeding occurs in groups, kleptoparasitism is common. Kleptoparasitism is most beneficial in procuring food during periods of food abundance and in large feeding aggregations. Immatures use kleptoparasitism as much as adults, but are attacked more often by adults than birds of similar age. Adults appear to benefit most from this behavior. The bold color patterns of adults may be an important signal influencing the formation of feeding groups. However, a video from Russia shows a juvenile Steller's sea eagle aggressively displacing an adult from food during a protracted battle.[33] Outside the breeding period, these eagles probably roost communally near their feeding sites. When salmon and trout are dying in winter after their summer spawning, feeding groups of Steller's sea eagles may mix with smaller golden eagles (Aquila chrysaetos) and white-tailed eagles to exploit this food source. This area is the only one in the golden eagle's nearly circumpolar range where they are extensively dependent on fish for prey.[34] Kleptoparasitism is sometimes recorded within the species. Occasionally, the smaller species may steal a fish away from the Steller's, especially if it is distracted by aggression from conspecifics, and both juvenile and adult Steller's may lose fish to the smaller species even face-to-face, especially a less assertive bird, such as immature Steller's. One video shows a golden eagle engaging an immature Steller's in a conflict and ultimately displacing it after maintaining a superior grip despite its smaller size.[35] In other cases, the Steller's have been photographed coming away with the prey after using its superior size to dominate, usually by baring down its mass and large bill over the smaller eagles.[36][37][38] In other cases, though, the three eagle species have been observed to feed in close proximity and seem to be outwardly indifferent to each other's presence. In inland areas, where golden, bald and white-tailed eagles compete over food sources which are not as abundant as these fish and, more importantly, nesting ranges, aggressive interspecies competition can be more common.[5] White-tailed eagles and golden eagles have even killed one another in Scotland, in cases of competition for abutting nesting ranges.[34] As in many sea and fish eagles, Steller's sea eagle may attempt to steal (and occasionally succeed in procuring) fish from osprey (Pandion haliaetus) where they coexist.[39] In one case, a cinereous vulture (Aegypius monachus), the largest living accipitrid, was observed to be pursued in flight and kleptoparasitized by a Steller's sea eagle.[40]

Reproduction[edit]

This eagle builds several aeries, being bulky constructions of twigs and sticks, at a height of up to 150 cm (59 in) and diameter of up to 250 cm (98 in). They usually place such nests high up on trees and rock at 15 to 20 m (49 to 66 ft) above the ground, sometimes in trees up to 45 m (148 ft). Alternate nests are usually built within 900 m (3,000 ft) of each other. In one case, two active nests were found to have been located within 100 m (330 ft).[5][13]

Courtship, which usually occurs between February and March, and reportedly simply consists of a soaring flight above the breeding area.[13] The Steller's sea eagle copulate on the nest after building it.[3] They lay their first greenish-white eggs around April to May. The eggs range from 78 to 85 mm (3.1 to 3.3 in) height and 57.5 to 64.5 mm (2.26 to 2.54 in) in width and weigh around 160 g (5.6 oz), being slightly larger than those of harpy eagles.[13] Clutches can contain from one to three eggs, with two being the average. Usually only one chick survives to adulthood, though in some cases as many as three will successfully fledge.[13] After an incubation period of around 39 – 45 days the chicks hatch, the helpless, whitish-down covered young are born. Incubation begins with the first hatching, which occurs in mid-May to late June. The eaglets fledge in August or early September. Adult plumage is attained at four years of age, but first breeding doesn't typically occur for another year or two.[3][13]

Eggs and very small nestlings can be preyed on by arboreal mammals, such as sables and ermine, and birds, usually corvids. Any of these small, clever nest predators rely on distraction and stealth to predate the eagle's nests and are killed if caught by either of the parents. Once it reaches roughly adult size in the fledging stage, few predators can threaten this species. In one case, a brown bear (Ursus arctos) was able to access a nest located on a rock formation and ate a fledging eaglet, though this is believed to be exceptional. Fully-grown fledgings in tree nests are probably invulnerable to predation, as there are no large mammalian carnivores which can climb trees in the species' range.[41] Due primarily to egg predation and nest collapses, only 45–67% of eggs are successfully reared to adulthood and up to 25% of nestlings may be lost.[5] However, once fully grown, the eagle has no natural predators.[24]

Conservation status[edit]

This species is classified as Vulnerable by the IUCN. They are legally protected, being classified as a National Treasure in Japan and mostly occurring in protected areas in Russia. However, many threats to their survival persist. These mainly include habitat alteration, industrial pollution and over-fishing, which in turn decreases their prey source. The current population is estimated at 5,000 and decreasing.[1] It was observed that recent heavy flooding, which may have been an effect of global climate change, caused almost complete nesting failure for the eagles nesting in Russian rivers due to completely hampering the ability of the parents to capture the fish essential to their nestlings' survival.[27] Persecution of the bird in Russia continues, due to its habit of stealing furbearers from trappers.[5] Due to a lack of other accessible prey in some areas, increasingly eagles on Hokkaido have moved inland and scavenged on sika deer carcasses left by hunters, exposing them to a risk of lead poisoning through ingestion of lead shot.[42]

In Kamchatka, 320 pairs have been recorded. An additional 89 nesting areas are not monitored. In the mountains of Koryakan and along the Bay of Penshina, over 1,200 pairs breed and at least 1,400 juveniles occur. About 500 pairs live in the Khabarovsk region of the Okhostsk coast, and 100 on the Shantar Islands. Another 600 pairs occur in the lower Amur. There are 280 pairs on Sakhalin Island and a few on the Kurile Islands. The total population is approximately 3,200 breeding pairs. Possibly up to 3,500 birds winter on Kamchatka, and another approximate 2,000 may occur on Hokkaido.[5]

Gallery[edit]

Footnotes[edit]

  1. ^ a b IUCN Red List 2013.
  2. ^ Etymology: Haliaeetus, New Latin for "sea-eagle". pelagicus, "of the open seas", from Ancient Greek pelagos, the ocean.
  3. ^ a b c d e f g h i j k l m n o p q Ferguson-Lees & Christie (2001). Raptors of the World. ISBN 0-7136-8026-1
  4. ^ "Steller's Sea Eagle – Los Angeles Zoo and Botanical Gardens". Lazoo.org. Retrieved 31 May 2011. 
  5. ^ a b c d e f g h i j k l (Russian) Species Synopsis Steller'S Sea Eagle. Fadr.msu.ru. Retrieved on 2012-08-21.
  6. ^ a b DEMENTEV, G., & Gladkov, N. A. (1954). Ptitsy Sovetskogo Soyuza [The Birds of Soviet Union]. Sovetskaya Nauka, Moskva, 5, 803.
  7. ^ a b c CRC Handbook of Avian Body Masses, 2nd Edition by John B. Dunning Jr. (Editor). CRC Press (2008), ISBN 978-1-4200-6444-5.
  8. ^ a b c d Saito, K. (2009). Lead poisoning of Steller’s Sea-Eagle (Haliaeetus pelagicus) and White-tailed Eagle (Haliaeetus albicilla) caused by the ingestion of lead bullets and slugs. Hokkaido Japan. In RT Watson, M. Fuller, M. Pokras, and WG Hunt (Eds.). Ingestion of Lead from Spent Ammunition: Implications for Wildlife and Humans. The Peregrine Fund, Boise, Idaho, USA.
  9. ^ a b Masterov, V. B. (2000). Postnatal development of Steller’s sea eagles in East Asia. In First symposium on Steller’s and white-tailed sea eagles in East Asia. Wild Bird Society of Japan, Tokyo (pp. 17-28).
  10. ^ Gamauf, A., Preleuthner, M. and Winkler, H. (1998). "Philippine Birds of Prey: Interrelations among habitat, morphology and behavior". The Auk 115 (3): 713–726. doi:10.2307/4089419. 
  11. ^ Steller's Sea Eagle. polarconservation.org. last update: April 21, 2009
  12. ^ Wood, Gerald (1983). The Guinness Book of Animal Facts and Feats. ISBN 978-0-85112-235-9. 
  13. ^ a b c d e f g h i j k l m n o p q r Eagles, Hawks and Falcons of the World by Leslie Brown & Dean Amadon. The Wellfleet Press (1986), ISBN 978-1555214722.
  14. ^ a b c Ladyguin, Alexander (2000). The morphology of the bill apparatus in the Steller’s Sea Eagle. First Symposium on Steller’s and White-tailed Sea Eagles in East Asia pp. 1–10; Ueta, M. & McGrady, M.J. (eds.) Wild Bird Society of Japan
  15. ^ Nakagawa, H. Steller's Sea Eagle. Bird Research News, 2009.2.24, pp. 2-3.
  16. ^ Tabaranza, Blas R., Jr. (2005-01-17). "The largest eagle in the world". Haribon Foundation. Retrieved 2012-09-23. 
  17. ^ Etymology: niger, Latin for "black".
  18. ^ Kaiser, M. (2010). A living specimen of the dark form of Steller’s Sea Eagle, Haliaeetus pelagicus ("niger") in captivity. J. Orn. Online. doi:10.1007/s10336-010-0580-2
  19. ^ Davies, E. (2010). Dark Steller's sea eagle solves 100 year debate. BBC Online. Accessed 26 October 2010.
  20. ^ "Steller's Sea-Eagle | San Diego Zoo Animals". Sandiegozoo.org. Retrieved 2013-10-07. 
  21. ^ Wink, Heidrich & Fentzloff 1996.
  22. ^ Whitfield, D. P., Duffy, K., McLeod, D. R., Evans, R. J., MacLennan, A. M., Reid, R., & Douse, A. (2009). Juvenile dispersal of white-tailed eagles in western Scotland. Journal of Raptor Research, 43(2), 110-120.
  23. ^ Mcgrady, M. J., Ueta, M., Potapov, E. R., Utekhina, I., Masterov, V., Ladyguine, A., & Seegar, W. S. (2003). Movements by juvenile and immature Steller's Sea Eagles Haliaeetus pelagicus tracked by satellite. Ibis, 145(2), 318-328.
  24. ^ a b c d "ADW: Haliaeetus pelagicus: Information". Animaldiversity.ummz.umich.edu. Retrieved 31 May 2011. 
  25. ^ "Steller's sea eagle videos, photos and facts – Haliaeetus pelagicus". ARKive. 17 April 2003. Retrieved 31 May 2011. 
  26. ^ a b c d e f g h Utekhina, I., Potapov, E., & McGRADY, M. J. (2000). Diet of the Steller’s Sea Eagle in the northern Sea of Okhotsk. In First Symposium on Steller’s and White-tailed Sea Eagles in East Asia. Tokyo, Japan: Wild Bird Society of Japan (pp. 71-92).
  27. ^ a b c Potapov, E., U. Irina, M. McGrady, and D. Rimlinger. 2010. (Abstract) Source-sink populations of the Steller's Sea Eagles (Haliaeetus pelagicus) in the northern part of the Sea of Okhotsk: ecological traps and their conservation implications. P. 32 in S. Gombobaatar, R. Watson, M. Curti, R. Yosef, E. Potapov, and M. Gilbert (eds.). Asian raptors: science and conservation for present and future: The proceedings of the 6th International Conference on Asian Raptors. Asian Rapto/r Research and Conservation Network, Mongolian Ornithological Society, and National University of Mongolia, Ulaanbaatar, Mongolia.
  28. ^ "Steller's sea eagle Haliaeetus pelagicus". The Peregrine Fund: Global Raptor Information Network. 2/12/2012. Retrieved 9 March 2013.  Check date values in: |date= (help)
  29. ^ "Steller's Sea Eagles, Steller's Sea Eagle Pictures, Steller's Sea Eagle Facts – National Geographic". Animals.nationalgeographic.com. Retrieved 31 May 2011. 
  30. ^ Ueta, M.; McGrady, M. J.; Nakagawa, H.; Sato, F.; Masterov, V. B. 2003. Seasonal change in habitat use in Steller's sea eagles. Oryx 37: 110-124.
  31. ^ Wood, The Guinness Book of Animal Facts and Feats. Sterling Pub Co Inc (1983), ISBN 978-0-85112-235-9.
  32. ^ BirdLife International (2003) BirdLife's online World Bird Database: the site for bird conservation. Version 2.0. Cambridge, UK: BirdLife International.
  33. ^ "Wild Russia- Eagles Battle Over Food". YouTube. Retrieved 2013-06-04. 
  34. ^ a b Jeff Watson (23 August 2010). The Golden Eagle. A&C Black. ISBN 978-1-4081-1420-9. Retrieved 23 October 2012. 
  35. ^ "Kamatchka:Golden Eagle fights Steller's Sea Eagle". Retrieved 2014-09-19. 
  36. ^ Steller's sea eagle photo – Haliaeetus pelagicus – G113853. ARKive. Retrieved on 2012-08-21.
  37. ^ The Secret Lives of Sea Eagles – National Wildlife Federation. Nwf.org (2011-10-26). Retrieved on 2012-08-21.
  38. ^ Photo Keywords : steller's sea eagle : White-tailed Eagle and Steller's Sea Eagle. Golden First Light (2012-06-21). Retrieved on 2012-08-21.
  39. ^ "Steller's Sea-Eagle". Woodland Park Zoo Seattle WA. Retrieved 2014-09-22. 
  40. ^ "Birds Korea- Bird News January 2006". Birds Korea. Retrieved 2013-05-22. 
  41. ^ McGrady, M.J., E. Potapov, and I. Utekhina. 1999. Brown Bear (Ursus arctos) feeds on Steller's Sea-eagle (Haliaeetus pelagicus) nestling. Journal of Raptor Research 33:342–343.
  42. ^ sheet (2011).
References
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