Overview

Brief Summary

Biology

In common with most other ducks, after mating the drake shoveler plays no further part in the incubation of the eggs and rearing of the ducklings. In April or May, the female constructs a nest on dry ground, often concealed in a grass tussock, and lines it with down from her own breast. There may be as many as 12 buff-coloured eggs and incubation takes 24 days. The female leads them away from the nest as soon as the clutch has hatched. The ducklings feed on small insects and other invertebrates, as well as plant seeds and buds. They can fly after about six weeks, and by the end of October, most British birds have migrated to southern Europe.
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Comprehensive Description

Description

The shoveler is named for its extraordinary oversized bill, which has a broad spatula-shaped tip. Both sexes have this feature, but the drake (male) shoveler, in his flamboyant breeding plumage, is easily distinguishable from the female. He has a bottle-green head (rather like the drake mallard), a white chest, chestnut flanks and black primary wing feathers and tail. The upper shoulder of the wing has a prominent sapphire blue flash. In the late summer, the drake loses his finery, and goes into 'eclipse plumage' after moulting. Both sexes then appear mottled brown, although the drake can be identified by a white streak just in front of the eye. Immature birds are similar in appearance to birds in this eclipse phase but look somewhat darker. Shovelers belong to the family Anatidae or dabbling ducks, and this describes their feeding behaviour exactly. Shovelers rarely 'up-end' like mallard and other surface-feeding ducks. However, they will dive if disturbed.
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Description of Anas clypeata

Het mannetje is opvallender gekleurd dan het vrouwtje
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Distribution

Global Range: (>2,500,000 square km (greater than 1,000,000 square miles)) BREEDING: Holarctic. In North America, primarily from Alaska east to Manitoba, south to California, New Mexico, Nebraska, western Iowa, locally eastward; also in Eurasia. NON-BREEDING: southwestern British Columbia, Arizona, east to Gulf Coast, coastal Georgia and South Carolina south to northern Colombia (rarely northern Venezuela), West Indies, and Hawaii, rarely north to north-central and northeastern U.S.; also Old World. In the U.S., the highest winter densities occur in the San Joaquin and Imperial valleys and the Clear Lake refuge in California, and the Bitter Lake refuge in New Mexico; winter abundance may vary greatly from year to year at a particular location (Root 1988).

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Geographic Range

Northern shovelers (Anas clypeata) has a very broad geographical range. They breed throughout Eurasia and western North America. They are also found in the Great Lakes region of the eastern United States. In winter various populations migrate south to specific locations, scattered throughout north-east Africa, India, Southern China and Japan to Mexico and southern North America. (Soothill and Whitehead, 1988)

Biogeographic Regions: nearctic (Native ); palearctic (Native ); oriental (Native )

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occurs (regularly, as a native taxon) in multiple nations

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National Distribution

Canada

Origin: Native

Regularity: Regularly occurring

Currently: Present

Confidence: Confident

Type of Residency: Year-round

United States

Origin: Native

Regularity: Regularly occurring

Currently: Present

Confidence: Confident

Type of Residency: Year-round

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Range

Shovelers are resident across most of Britain although those in the north migrate south to avoid harsh northern winters. The largest UK populations are found in East Anglia and central England. The birds range across most of southern and central Europe, Finland and Russia including Siberia, and are also found in central and western parts of the USA and Canada. Most of the birds that breed in northern latitudes migrate south during the winter.
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Physical Description

Morphology

Physical Description

Males weigh 17 to 38 ounces(470 to 1000g) and their wingspans are usually around 31 inches(227 to 251mm). Females are 17 to 28 ounces(470 to 800g). Northern Shovelers are sexually dimorphic. The males head, neck, and speculum are iridescent green, their chests are white, and the remaining underparts are a bright chestnut. The females are mainly a pattern of buffs and browns. Both sexes have pale blue inner forewings and orange-yellow legs and feet. The most distinctive feature is their large spatulate bill. It is twice as wide at the tip than it is at the base. This uniquely shaped bill gives rise to Northern Shovelers also being called "spoonbills". The ducklings hatch with a typical duckbill that enlarges as the duckling matures. (Goodes and Boyer, 1986; Todd, 1979)

Range mass: 470 to 1000 g.

Average basal metabolic rate: 3.9209 W.

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Size

Length: 48 cm

Weight: 636 grams

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Ecology

Habitat

Habitat and Ecology

Habitat and Ecology
Behaviour This species is highly migratory although it may be present all year round in parts of Europe1. It arrives on the breeding grounds from March3 where it breeds in solitary pairs or loose groups1 in the northern spring (chiefly from mid-April to June)3. Males undergo a post-breeding moult migration from early-May to early-June (females moulting one month later) during which they are flightless for 3-4 weeks5. The autumn migration chiefly occurs between September and October (western Europe), during which the species is likely to travel on a broad front (e.g. across Arabia and into Africa)5. The species is usually found in pairs or small parties3, 4 although it often congregates when feeding2 (flocks of 20 or 30 to several hundred individuals occur in favoured areas in Africa6 and large concentrations form at stop-over sites on migration)3. The species forages diurnally and roosts communally at night6. Habitat It inhabits permanent8 shallow freshwater wetlands1 from sea level up to 2,900 m (Ethiopia)6, preferred sites being those surrounded by dense stands of reeds or other emergent vegetation whilst being free of overhanging trees or fringing forest8. Copious submerged aquatic vegetation sheltering abundant planktonic invertebrates is a valuable habitat characteristic4. Suitable habitats include well-vegetated lakes and marshes and with muddy shores and substrates in open country1, 2, 3 (e.g. grasslands)4, 6, as well as oxbow lakes, channels and swamps (former U.S.S.R.)7. It also frequents artificial waters bordered by lush grassland8 such as sewage farms, rice-fields2 and fish ponds11. In the winter it can be found on coastal brackish lagoons1, 2, 3, tidal muflats1, 2, estuaries3, coastal shorelines4, fresh and brackish estuarine marshes4, inland seas and brackish or saline inland waters8, occasionally occurring (briefly) on marine waters during migration3, 8 (although it generally avoids very saline habitats)4. Diet Its diet consists of small aquatic invertebrates1 such as adult and larval insects1 (e.g. caddisfly larvae, damselfly and dragonfly nymphs, adult beetles, bugs4 and flies6), molluscs1, planktonic crustaceans8, the seeds1 of emergent and aquatic plants2 (e.g. bulrushes and waterweeds)4, annelids, amphibian spawn, tadpoles, spiders, fish and the vegetative parts of aquatic plants6 (e.g. duckweeds)4, 6. Breeding site The nest is a scrape2 or depression1 on the ground in tall grass, among hummocks, in the open7 or (rarely) in bulrush marshes2. Usually the species nests close to water but if grass cover is unavailable in the wetland site it may also nest far away from water under bushes7, in hayfields or in meadows2. Although it is not a colonial species, several pairs may nest in close proximity3. Management information A study in the Czech Republic found that fish ponds with a fish stock density of less than 400 kg/ha, water transparency of more than 50 cm, mixed fish stocks (e.g. tench and pike or perch) rather than monospecific stocks (e.g. of carp), and systems that include ponds with fish fry (to provide areas with low fish competition and high invertebrate availability) are more successful in supporting breeding pairs of this species11. The cyclical removal of adult fish from an artificial waterbody (gravel pit) in the UK attracted nesting pairs to the area by causing an increase in invertebrate food availability and an increase in the growth of submerged aquatic macrophytes25. The removed fish (dead or alive) were sold to generate funds25.

Systems
  • Terrestrial
  • Freshwater
  • Marine
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Habitat

During the breeding season, Northern Shovelers are found in shallow pools and marshes that have good cover and dry areas nearby for nesting. In the winter they can be found near freshwater marshes, swamps, and flooded areas. (Johnsgard 1965.)

Terrestrial Biomes: savanna or grassland ; forest

Aquatic Biomes: lakes and ponds

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Depth range based on 1 specimen in 1 taxon.
Water temperature and chemistry ranges based on 1 sample.

Environmental ranges
  Depth range (m): 0 - 0
  Temperature range (°C): 10.275 - 10.275
  Nitrate (umol/L): 7.182 - 7.182
  Salinity (PPS): 33.744 - 33.744
  Oxygen (ml/l): 6.451 - 6.451
  Phosphate (umol/l): 0.514 - 0.514
  Silicate (umol/l): 4.619 - 4.619
 
Note: this information has not been validated. Check this *note*. Your feedback is most welcome.
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Comments: BREEDING: Shallow, often muddy, fresh-water areas with surrounding cover. Ponds, marshes, sloughs, and creeks. Nests near shallow freshwater lake, pond, marsh, etc. Nests on the ground, usually near edge of water. The nest is a hollow lined with plant material and down. NON-BREEDING: In migration and winter in both freshwater and brackish habitats, and in cultivated fields (not typical) (AOU 1983).

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Habitat

Shovelers prefer areas of shallow fresh water surrounded by rich vegetation, although they can be found on fens and marshes with plenty of open water.
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Migration

Non-Migrant: No. All populations of this species make significant seasonal migrations.

Locally Migrant: No. No populations of this species make local extended movements (generally less than 200 km) at particular times of the year (e.g., to breeding or wintering grounds, to hibernation sites).

Locally Migrant: Yes. At least some populations of this species make annual migrations of over 200 km.

Small flocks migrate northward in spring, break up into pairs or small groups upon arrival in nesting areas. Flocks start migrating southward in late August or early September. (departs far north July-August). Present in northern South America mostly October-March (Hilty and Brown 1986).

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Trophic Strategy

Food Habits

A Northern Shoveler feeds mainly by drawing water into its bill and then pumping it out through the sides with their tongue, filtering out minute food particles with long comb-like lamellae that line the edge of the bill. The particles mainly consist of tiny crustaceans, molluscs, insects, and their larvae as well as seeds and pieces of leaves and stems of plants. In addition to the food particles they also eat water beetles, small minnows, and snails. Social feeding is common. The shovelers are drawn to feeding areas by other birds feeding in an area. Shovelers take advantage of the food particles churned to the surface by the other birds swimming or wading in the area. Single birds may swim in a tight circle to create a whirlpool to cause food to come to the surface. Shovelers are also known to upend or dabble, usually for lengthier periods than other surface feeders, and also dive using their wings to swim underwater in shallow marshes. (Gooders and Boyer, 1986, Johnsgard, 1969, Todd, 1979)

Animal Foods: terrestrial non-insect arthropods; mollusks; aquatic or marine worms; aquatic crustaceans

Foraging Behavior: filter-feeding

Primary Diet: carnivore (Insectivore , Eats non-insect arthropods, Molluscivore )

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Comments: Opportunistic forager. Eats seeds of sedges, bulrushes, saw grass, pondweeds, smartweeds, algae, duckweeds, etc; also mollusks, aquatic insects, and crustaceans. In Manitoba, males and females ate primarily aquatic invertebrates during prelaying and laying periods (Ankney and Afteon 1988). Aquatic invertebrates (e.g., water boatmen) may dominate winter diet in some areas. Usually dabbles at water surface.

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General Ecology

Large concentrations seen at migration staging areas. Usually feeds in pairs or small groups.

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Life History and Behavior

Life Expectancy

Lifespan/Longevity

Average lifespan

Status: wild:
223 months.

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Lifespan, longevity, and ageing

Maximum longevity: 20.4 years
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Reproduction

Reproduction

Breeding usually takes place from April until June. Nests are made on dry land close to fresh water and they are built of grasses and lined with down feathers. The female builds the nest by forming a neat cup by twisting her body on the ground. She lays between 9 and 11 eggs. The eggs are olive colored and 52x37mm. Incubation by the female alone begins immediately after all the eggs are laid and can last 23 to 25 days. (Dobkin, Ehrlich and Wheye, 1988.) (Soothill and Whitehead, 1978.)

Range eggs per season: 9 to 11.

Average time to hatching: 23 days.

Average eggs per season: 11.

The male loses interest soon after incubation starts. The ducklings are born precocial and start following the female almost immediately. Feeding practices and locations are learned during this time. They young can fly after 40 to 45 days and are then independent.

Parental Investment: precocial

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Breeding begins in late March in the south to early June in the north. Clutch size: 6-14 (usually 10-12). Incubation: 23-25 days, by female (Terres 1980). Young are tended by female, independent in about 6-7 weeks (Harrison 1978). Clutch size may be limited by lipid reserves rather than by protein acquisition (Ankney and Afteon 1988).

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Molecular Biology and Genetics

Molecular Biology

Barcode data: Anas clypeata

The following is a representative barcode sequence, the centroid of all available sequences for this species. 

 
There are 18 barcode sequences available from BOLD and GenBank.  Below is a sequence of the barcode region Cytochrome oxidase subunit 1 (COI or COX1) from a member of the species.  See the BOLD taxonomy browser for more complete information about this specimen and other sequences.
 
BON465-07|NHMO-BC468|Anas clypeata| ---------------------------CAAGACATTGGCACTCTATACCTTATCTTCGGGGCATGAGCCGGAATAATCGGCACAGCACTG---AGCCTACTAATCCGCGCAGAACTTGGTCAACCAGGGACTCTCCTGGGCGAT---GACCAAATTTACAACGTGATCGTCACCGCTCACGCCTTTGTAATAATCTTCTTCATAGTAATACCCATCATAATTGGAGGATTTGGCAACTGATTGGTCCCCCTGATA---ATCGGTGCCCCAGACATAGCATTCCCACGAATAAATAACATAAGCTTCTGACTCCTTCCACCATCATTCCTTCTGCTACTCGCCTCATCTACCGTAGAAGCTGGCGCTGGTACAGGTTGAACCGTGTACCCACCCCTAGCAGGCAATCTGGCCCACGCTGGGGCCTCAGTAGACCTG---GCCATTTTCTCGCTCCACCTAGCCGGTGTTTCCTCCATCCTCGGGGCTATTAACTTCATTACCACAGCCATCAACATAAAACCCCCTGCACTCTCACAGTACCAAACCCCACTTTTCGTCTGATCGGTCCTAATTACCGCCATCCTGCTCCTCCTGTCACTTCCTGTCCTTGCCGCC---GGCATCACAATGCTACTAACCGACCGAAACCTAAACACCACATTCTTCGACCCTGCTGGAGGGGGAGACCCGATCCTGTACCAACACCTATTTTGATTCTTCGGCCACCCAGAAGTCTACATCTTAATTCTTCCAGGATTTGGAATT---------------------------------------------------------------------------------------------------------------------------------------- 
-- end --

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Statistics of barcoding coverage: Anas clypeata

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 18
Species: 23
Species With Barcodes: 1

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Conservation

Conservation Status

IUCN Red List Assessment


Red List Category
LC
Least Concern

Red List Criteria

Version
3.1

Year Assessed
2009

Assessor/s
BirdLife International

Reviewer/s
Bird, J., Butchart, S.

Contributor/s

Justification
This species has an extremely large range, and hence does not approach the thresholds for Vulnerable under the range size criterion (Extent of Occurrence <20,000 km2 combined with a declining or fluctuating range size, habitat extent/quality, or population size and a small number of locations or severe fragmentation). Despite the fact that the population trend appears to be decreasing, the decline is not believed to be sufficiently rapid to approach the thresholds for Vulnerable under the population trend criterion (>30% decline over ten years or three generations). The population size is extremely large, and hence does not approach the thresholds for Vulnerable under the population size criterion (<10,000 mature individuals with a continuing decline estimated to be >10% in ten years or three generations, or with a specified population structure). For these reasons the species is evaluated as Least Concern.

History
  • 2008
    Least Concern
  • 2004
    Least Concern
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Conservation Status

The population in North America appears to be increasing. (Todd, 1979,) (Wcmc.org, July 10, 2000.) This species is protected under the U.S. Migratory Bird Act.

US Migratory Bird Act: protected

US Federal List: no special status

CITES: no special status

State of Michigan List: no special status

IUCN Red List of Threatened Species: least concern

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Status in Egypt

Regular passage visitor and winter visitor.

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IUCN

Least Concern.

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National NatureServe Conservation Status

Canada

Rounded National Status Rank: N4N,N5B : N4N: Apparently Secure - Nonbreeding, N5B: Secure - Breeding

United States

Rounded National Status Rank: N5B,N5N : N5B: Secure - Breeding, N5N: Secure - Nonbreeding

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NatureServe Conservation Status

Rounded Global Status Rank: G5 - Secure

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Status

Receives general protection under the Wildlife and Countryside Act (as amended) in the UK. Included in the Birds of Conservation Concern Amber List (medium conservation concern).
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Threats

Threats

Major Threats
The species is threatened by habitat loss in Britain and Ireland2, is occasionally killed by collisions with power transmission lines23, and suffers from nest predation by American mink Neovison vison17, 19, 24. It is susceptible to avian influenza10, 21 and avian botulism18 so may be threatened by future outbreaks of these diseases. The species may suffer from reproductive impairment as a result of selenium (Se) accumulation in liver tissues (selenium contained in sub-surface agricultural drain-water used for wetland management in California led to bioaccumulation of the element in the food chain)22. The species suffers mortality as a result of lead shot ingestion (Camargue, France15 and Spain20). Utilisation The species is a favoured quarry species throughout most of the world, but is rarely taken in large numbers2. It is hunted for sport in North America9, Denmark12 and the Po delta, Italy13, and is hunted commercially and recreationally in Iran14. The eggs of this species used to be (and possibly still are) harvested in Iceland16.
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Threats

The shoveler does not appear to be a threatened species, although the north-west European population is believed to have been in decline for some years. Between the late 1960s and the early 1990s, there was a 39% reduction in the number of 10 kilometre squares where breeding was recorded in the UK. Some of this decline is thought to be through the loss of wetland in several parts of the country, and the British breeding population (currently estimated at about 1000 pairs) is concentrated at a relatively small number of sites. The species receives general protection under the Wildlife and Countryside Act (as amended), the Wildlife (Northern Ireland) Order and the EC Birds Directive.
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Management

Restoration Potential: No difference between natural and restored wetlands in pair counts over two years in Iowa (Dubowy 1996).

Management Requirements: See Barker et al. (1990) for information on the effects of different livestock grazing systems on nesting success in North Dakota.

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Conservation

The single most important UK site for the shoveler is the Ouse Washes in Cambridgeshire, thought to support over 150 pairs, some 2% of the international population. The Ouse Washes hold significant numbers of many breeding waterfowl and have been designated as a Special Protection Area (SPA). This site is specially managed for wildfowl, being allowed to flood during the winter, and grazed to encourage suitable conditions for breeding birds. In addition to the breeding population, shoveler numbers are swelled in winter by migratory birds from northern Europe, Russia and the Ukraine, and (possibly) Iceland. Shoveler migration is a complex affair, with British birds leaving for southern Europe and northern European birds arriving to overwinter. All this makes assessing population sizes rather difficult, so the current estimate of 10,000 UK birds may be less than the actual number. In addition to the designated SPA for breeding shovelers, there are 26 non-breeding SPAs, where the birds' importance qualifies the sites for special protection.
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Relevance to Humans and Ecosystems

Benefits

Economic Importance for Humans: Negative

None found.

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Economic Importance for Humans: Positive

Northern Shovelers are a game bird. Hunters often shoot them due to their resemblance to mallards. They are often referred to as "neighbor's mallards," because some hunters give them to their neighbors and keep the more tasty mallards for themselves. (Todd, 1979.)

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Wikipedia

Northern Shoveler

The Northern Shoveler (play /ˈʃʌvələr/; Anas clypeata), Northern Shoveller in British English, sometimes known simply as the Shoveler, is a common and widespread duck. It breeds in northern areas of Europe and Asia and across most of North America,[2] wintering in southern Europe, Africa, the Indian subcontinent, Southeast Asia, and Central and northern South America. It is a rare vagrant to Australia. In North America, it breeds along the southern edge of Hudson Bay and west of this body of water, and as far south as the Great Lakes west to Colorado, Nevada, and Oregon.[3][4]

The Northern Shoveler is sometimes referred to by hunters as the "spoony". Other disparaging names, as compared to the mallard, are the "smiling mallard" and the "Poor Man's Mallard".[5]

The Northern Shoveler is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) applies.[6] The conservation status of this bird is Least Concern.[1]

Contents

Taxonomy

This species was described by Linnaeus in his Systema naturae in 1758 under its current scientific name.[7] Usually placed in Anas like most dabbling ducks, it stands well apart from such species as the Mallard and together with the other shovelers and their relatives forms a "blue-winged" group that may warrant separation as genus Spatula.

No living subspecies are accepted today. Fossil bones of a very similar duck have been found in Early Pleistocene deposits at Dursunlu, Turkey. It is unresolved, however, how these birds were related to the Northern Shoveler of today; i.e., whether the differences noted were due to being a related species or paleosubspecies, or attributable to individual variation.[8]

Description

Female stretching after bathing in Kolkata

This species is unmistakable in the northern hemisphere due to its large spatulate bill. The breeding drake has an iridescent dark green head,[5] white breast and chestnut belly and flanks. In flight, pale blue forewing feathers are revealed, separated from the green speculum by a white border. In early fall the male will have a white crescent on each side of the face.[4] In non-breeding (eclipse) plumage, the drake resembles the female.

The female is a drab mottled brown like other dabblers,[5] with plumage much like a female Mallard, but easily distinguished by the long broad bill, which is gray tinged with orange on cutting edge and lower mandible.[4] The female's forewing is gray.

They are 19 inches (48 cm) long and have a wingspan of 30 inches (76 cm) with a weight of 600 grams (1.3 lb).[3]

Behavior

In flight

Northern Shovelers feed by dabbling for plant food, often by swinging its bill from side to side and using the bill to strain food from the water. They use their highly specialized bill (from which their name is derived) to forage for aquatic invertebrates – a carnivorous diet. Their wide-flat bill is equipped with well-developed lamellae – small, comb-like structures on the edge of the bill that act like sieves, allowing the birds to skim crustaceans and plankton from the water's surface. This adaptation, more specialized in shovelers, gives them an advantage over other puddle ducks, with which they do not have to compete for food resources during most of the year. Thus, mud-bottomed marshes rich in invertebrate life are their habitat of choices.[5]

The shoveler prefers to nest in grassy areas away from open water. Their nest is a shallow depression on the ground, lined with plant material and down. Hens typically lay about nine eggs. The drakes are very territorial during breeding season and will defend their territory and partners from competing males. Drakes also engage in elaborate courtship behaviors, both on the water and in the air; it is not uncommon for a dozen or more males to pursue a single hen. Despite their stout appearance, shovelers are nimble fliers.[5]

This is a fairly quiet species. The male has a clunking call, whereas the female has a Mallard-like quack.

Habitat and range

This is a bird of open wetlands, such as wet grassland or marshes with some emergent vegetation.

This bird winters in southern Europe, Africa, the Indian Subcontinent, northern South America, and the Malay Archipelago.[2] Those wintering in the Indian Subcontinent make the taxing journey over the Himalayas, often taking a break in wetlands just south of the Himalaya before continuing further south to warmer regions. In North America it winters south of a line from Washington to Idaho and from New Mexico east to Kentucky, also along the Eastern Seaboard as far north as Massachusetts.[3][4] In the British Isles, home to more than 20% of the North Western European population, it is best known as a winter visitor, although it is more frequently seen in southern and eastern England, especially around the Ouse Washes, the Humber and the North Kent Marshes, and in much smaller numbers in Scotland and western parts of England. In winter, breeding birds move south, and are replaced by an influx of continental birds from further north. It breeds across most of Ireland, but in very small numbers.

This dabbling duck is strongly migratory and winters further south than its breeding range (so far so that there have been four reports in Australia).[citation needed] It is not as gregarious as some dabbling ducks outside the breeding season and tends to form only small flocks. Among North America's duck species, northern shovelers trail only mallards and blue-winged teal in overall abundance. Their populations have been healthy since the 1960s, and have soared in recent years to more than 4 million birds (2011), most likely because of favorable breeding, migration, and wintering habitat conditions.[5]

Gallery

Footnotes

  1. ^ a b IUCN (2009)
  2. ^ a b Clements, J. (2007)
  3. ^ a b c Floyd, T. (2008)
  4. ^ a b c d Dunn, J. (2006)
  5. ^ a b c d e f Ducks Unlimited, March/April 2010
  6. ^ AEWA
  7. ^ Linnaeus, C. (1758)
  8. ^ Louchart, A et al. (1998)

References

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Names and Taxonomy

Taxonomy

Comments: See Livezey (1991) for a phylogenetic analysis and classification (supergenera, subgenera, infragenera, etc.) of dabbling ducks based on comparative morphology; one conclusion was that the four species of shovelers are monophyletic.

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