Comprehensive Description
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The species G. bicalcaratus has originally been described from Nosy Boraha, and this name afterwards been applied widely to populations of brown and inconspicuously coloured Pandanus frogs. Genetic data indicate that most of these populations are strongly differentiated from Nosy Boraha specimens. Furthermore, two clearly distinct species occur syntopically at Nosy Boraha.
This species can be distinguished from other species of the Pandanusicola subgenus by differences in tympanum diameter and coloration. M. liber has larger hands and feet (Glaw et al. 1994).
Taken with permission from Glaw and Vences (2007).
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