The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 1996. database, NODC Taxonomic Code
  • 2004. database, Amphibian Species of the World: an Online Reference
  • Allmon, W. 1991. A plot study of forest floor litter frogs, Centeral Amazon, Brazil. Journal of Tropical Ecology, 7: 503-522.
  • Cekalovic K. 1985. Catálogo de Opiliones de Chile (Arachnida). Boletín de la Sociedad de Biología de Concepción (Chile) 56: 7-29. Cakalovic K. 1976. Catálogo de los Arachnida: Scorpiones, Pseudoscorpiones, Opiliones, Acari, Aranea y Solifugae de la XII Región de Chile, incluyendo la Antártica chilena (Chile). Gayana Zool. (Chile) 37: 1-108.
  • Clemente-Carvalho, R., L. Monteiro, V. Bonato, H. Rocha, G. Pereira, D. Oliveira. 2008. Geographic variation in cranial shape in the pumpkin toadlet (Brachycephalus ephippium): a geometric analysis. Journal of Herpetology, 44/4: 176-185.
  • Darst, C., D. Cannatella. 2004. Novel relationships among hyloid frogs inferred from 12s and 16s mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 31/2: 462-475.
  • Frost, D., T. Grant, J. Faivovich, R. Bain, A. Haas, C. Haddad. 2006. The amphibian tree of life. Bulletin of the America Museum of Natural History, 297: 1-371.
  • Frost, Darrel R. 1997. Amphibian Species of the World, A Taxonomic and Geographic Reference. Second edition
  • Frost, Darrel R. 1997. Amphibian Species of the World, A Taxonomic and Geographic Reference. Second edition
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Giaretta, A., K. Facure, J. Sawaya, J. Meyer, N. Chemin. 2006. Diversity and abundance of litter frogs in a montane forest of southeastern Brazil: seasonal and altitudinal changes. Biotropica, 31/4: 669-674.
  • Heyer, W., A. Rand, C. Goncalvez da Cruz, O. Peixoto. 1988. Decimations, extinctions, and colonizations of frog populations in southeast Brazil and their evolutionary implications. Biotropica, 20/3: 230-235.
  • International Union of Conservation of Nature and Natural Resources. 2010. "Brachycephalus ephippium" (On-line). Red List. Accessed February 06, 2011 at
  • Kaplan, M. 2002. Histology of the anteroventral part of the breast-shoulder apparatus of Brachycephalus ephippium with comments on the validity of the genus Psyllophryne. Amphibia-Reptilia, 23/2: 225-227.
  • Orrell T. (custodian) (2015). ITIS Global: The Integrated Taxonomic Information System (version Mar 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Pires, O., A. Sebben, E. Schwartz, R. Morales, C. Block, C. Schwartz. 2005. Further report of the occurrence of tetrodotoxin and new analogues in the Anuran family Brachycephalidae. Toxicon, 45/1: 73-79.
  • Pires, O., A. Sebben, E. Schwartz, S. Largura, C. Bloch, R. Morales, C. Schwartz. 2002. Occurrence of tetrodotoxin and its analogues in the Brazilian frog Brachycephalus ephippium (Anura: Brachycephalidae). Toxin, 40/6: 761-766.
  • Pombal, J. 2003. Pumpkin toadlet, Brachycephalus ephippium. Pp. 179-182 in M Hutchins, W Duellman, N Schlager, eds. Grzimek's Animal Life Encyclopedia, Vol. 6, 2 Edition. Farmington Hills, Michigan: Gale Group.
  • Pombal, J., E. Wistuba, M. Bornschein. 1998. A new species of Brachycephalid (anura) from the Atlantic Rain Forest of Brazil. Journal of Herpetology, 32/1: 70-74.
  • Pombal, J., I. Sazima, C. Haddad. 1994. Breeding behavior of the pumpkin toadlet, Brachycephalus ephippium. Journal of Herpetology, 28/4: 516-519.
  • Sebben, A., C. Schwartz, D. Valente, E. Mendes. 1986. A tetrodotoxin-like substance found in the Brazilian frog Brachycephalus ephippium. Toxicon, 40/6: 799-806.
  • Toledo, L., R. Ribeiro, C. Haddad. 2007. Anurans as prey: An exploratory analysis and size relationships between predators and their prey. Journal of Zoology, 271: 170-177.
  • Tuberville, T., V. Burke. 1994. Do flag markers attract turtle nest predators?. Journal of Herpetology, 28/4: 514-516.


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