The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Kolbe H.J. 1914. Wissenschaftliche Ergebnisse der deutschen Zentral-Afrika-Expedition 1907-1908 unter Führung Adolf Friedrichs, Herzog zu Mecklemburg. Band V: Zoologie III. Lieferung 3. Scarabaeiden. Klinkhardt & Biermann. Leipzig 5(3):261-416
  • Schoolmeesters P. (2013). Scarabs: World Scarabaeidae Database (version Aug 2012). In: Species 2000 & ITIS Catalogue of Life, 11th March 2013 (Roskov Y., Kunze T., Paglinawan L., Orrell T., Nicolson D., Culham A., Bailly N., Kirk P., Bourgoin T., Baillargeon G., Hernandez F., De Wever A., eds). Digital resource at Species 2000: Reading, UK.


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