The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 2001. website, U. S. Fish and Wildlife Service Endangered Species Program
  • 2001. website, Amphibian Species of the World: An online reference
  • 2004. database, Amphibian Species of the World: an Online Reference
  • Crump, M.L. (1988). ''Aggression in Harlequin Frogs: male-male competition and a possible conflict of interest between the sexes.'' Animal Behaviour, 36(4), 1064-1077.
  • Daly, J. W. (2004). ''Marine toxins and nonmarine toxins: convergence or symbiotic organisms?'' Journal of Natural Products, 67, 1211-1215.
  • Dunn, E. R. (1933). ''Amphibians and reptiles from El Valle de Anton, Panama.'' Occasional Papers of the Boston Society of Natural History, 8, 65-79.
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Karraker, N. E., Richards, C. L., and Ross, H. L. (2006). ''Reproductive ecology of Atelopus zeteki and comparisons to other members of the genus.'' Herpetological Review, 37(3), 284-288.
  • La Marca, E., Lötters, S., Puschendorf, R., Ibáñez, R., Rueda-Almonacid, J. V., Schulte, R., Marty, C., Castro, F., Manzanilla-Puppo, J., García-Pérez, J. E., Bolaños, F., Chaves, G., Pounds, J. A., Toral, E., and Young, B. E. (2005). ''Catastrophic population declines and extinctions in neotropical harlequin frogs (Bufonidae: Atelopus).'' Biotropica, 37(2), 190-201.
  • Lindquist, E. D. and Hetherington, T. E. (1998). ''Semaphoring in an earless frog: the origin of a novel visual signal.'' Animal Cognition, 1, 83-87.
  • Lindquist, E. D. and Hetherington, T. E. (1998). ''Tadpoles and juveniles of the Panamanian Golden Frog, Atelopus zeteki (Bufonidae), with information on development of coloration and patterning.'' Herpetologica, 54(3), 370-376.
  • Lindquist, E. D., and Hetherington, T. E. (1996). ''Field studies on visual and acoustic signaling in the ''earless'' Panamanian Golden Frog, Atelopus zeteki.'' Journal of Herpetology, 30(3), 347-354.
  • Lips, K. R., Brem, F., Brenes, R., Reeve, J. D., Alford, R. A., Voyles, J., Carey, C., Livo, L., Pessier, A. P., and Collins, J. P. (2006). ''Emerging infectious disease and the loss of biodiversity in a Neotropical amphibian community.'' Proceedings of the National Academy of Sciences, 103(9), 3165-3170.
  • Orrell T. (custodian) (2013). ITIS Global: The Integrated Taxonomic Information System (version Sept 12). In: Species 2000 & ITIS Catalogue of Life, 11th March 2013 (Roskov Y., Kunze T., Paglinawan L., Orrell T., Nicolson D., Culham A., Bailly N., Kirk P., Bourgoin T., Baillargeon G., Hernandez F., De Wever A., eds). Digital resource at Species 2000: Reading, UK.
  • Paratype: Dunn, E. R. 1933. Occ. Pap. Boston Soc. Nat. Hist. 8: 71.
  • Poole, V. (2006). Golden Frog Husbandry Manual, 2nd ed. [Internet PDF]. Denver: Denver Zoological Foundation's Project Golden Frog. Retrieved 2 October 2007, from
  • Richards-Zawacki, C. L. (2010). ''Thermoregulatory behaviour affects prevalence of chytrid fungal infection in a wild population of Panamanian golden frogs.'' Proceedings of the Royal Society B, 277, 519-528.
  • Savage, J. M. (2002). The Amphibians and Reptiles of Costa Rica. University of Chicago Press, Chicago and London.
  • Savage, J.M. (1972). ''The harlequin frogs, genus Atelopus, of Costa Rica and western Panama.'' Herpetologica, 28(2), 77-94.
  • Stuart, S., Hoffmann, M., Chanson, J., Cox, N., Berridge, R., Ramani, P., and Young, B. (eds) (2008). Threatened Amphibians of the World. Lynx Edicions, IUCN, and Conservation International, Barcelona, Spain; Gland, Switzerland; and Arlington, Virginia, USA.
  • Yotsu-Yamashita, M., Kim, Y.H., Dudley, Jr., S.C., Choudhary, G., Pfahnl, A., Oshima, Y., and Daly, J.W. (2004). ''The structure of zetekitoxin AB, a saxitoxin analog from the Panamanian golden frog Atelopus zeteki: a potent sodium-channel blocker.'' Proceedings of the National Academy of Sciences of the United States of America, 101(13), 4346-4351.


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