IUCN threat status:

Critically Endangered (CR)

Comprehensive Description

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Description

Males range from 29.9 to 37.1 mm SVL while females range from 35.0-46.0 mm SVL. The head is wide and long, with a snout that is acuminate in dorsal view and protrudes beyond the lower jaw in profile. The pretympanic and tympanic areas are covered with warts. The tympanic membrane, tympanic annulus, and stapes are absent. A glandular, slightly raised postorbital crest is present, but a postocular crest is lacking. The skin is generally smooth dorsally with some individuals having scattered spiculae or warts. The throat, chest, belly, and undersides of hindlimbs are rugose or areolate. Warts are present lateral to cloacal opening.

The forearms are relatively short. Webbing is absent on the fingers, and they lack lateral fringes. The fingers have digital tips with rounded pads. The third finger is the longest, followed by the fourth, second and first. Webbing is present in between the toes, with relative toe length IV>V=III>II>I and distinct digital pads. Many tubercles are present on the hand: palmar tubercle is round, supernumerary palmar tubercles are distinct, thenar and subarticular tubercles are distinct. On the foot, the inner metatarsal tubercle is oval, the outer metatarsal tubercle is round and raised, and the supernumerary plantar and subarticular tubercles are conspicuous. Males have short, robust forearms compared to females, and males also have keratinized nuptial pads on the dorsal and inner surfaces of the thumb and inner side of the proximal phalanx of Finger II. Males also have vocal slits.

The dorsum and dorsal surfaces of limbs and flanks are uniform reddish orange, orange, or dark brownish orange. Dorsal surfaces of fingers and toes are yellowish orange with brown areas. White spiculae are visible on the flanks, forearms and dorsal surfaces of the thighs. The black iris lacks a pupillary ring.

Tadpoles are unknown.

The specific name was given in dedication to the Guanujo, a Quechua people inhabiting the area where this species was once abundant, and the local town of the same name The species is known locally as "puca sapo", meaning "red toad".

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