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BiologyInhabits clear water streams, running over rocky, pebbles and sand bottom at low altitudes ranging from 188 to 258 m, with slow to moderate water current and less than 1 m deep (Ref. 81202). Stomach contents of three specimens include filamentous algae, fragments of vascular plants and seeds, insects and other unidentified arthropods, and organic debris (Ref. 81202).