IUCN threat status:

Vulnerable (VU)

Comprehensive Description

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The male body length is about 48 mm. The pupil is vertical. The tympanum is hidden. The maxillary teeth are developed(Fei 1999). The dorsum is relatively smooth(Wu et. al 1993) and yellowish brown with many, evenly scattered black spots. The head and the dorsal sides of the limbs are full of tubercles. A faint triangular mark is present between the eyes. The tibiotarsal articulation extends as far forward as the corner of the eye. The tips of the fingers and toes are rounded. The long toes are fringed and have rudimentary webbing(Fei 1999). The sides of the body are lighter in color than the back, with black spots decreasing in size ventrally. The hind legs are about 1.6 times the body length, and the heels overlap when they are flexed. Both the axilla and the back of the thigh have a light glandular spot. The backs of the limbs have irregular dark markings. The ventrum is smooth and grayish(Wu et. al 1993) with little or no spotting. The edge of the lips has black-brown horizontal stripes that are relatively distinct (Fei 1999). The male has black nuptial spines, positioned in three longitudinal rows, on the first and second finger. In addition, the edges of the jaws have small black spines (Wu et. al 1993). A relatively long chest gland, also possessing black spines, is positioned sideways on either side of the chest. The spines on the finger are large and thick and about two times the size of the spines on the chest. No vocal sac is present (Fei 1999). Eggs are 3.5 mm in diameter, milky white, and attached to the undersides of submerged rocks. The tadpoles are 65 mm in total body length and about 28 mm in head length. The back is brown and the tail muscle is a light yellowish brown with black-brown colored spots of varying sizes (Fei 1999). The lips are fringed with short, thick papillae, with pigmentation on their bases or extending to their tips. The median gap in papillae on the upper lip is about twice the width of the flanking papillae. The lower lip usually has two rows of compressed inframarginal papillae, most of which have small labial teeth (Wu et. al 1993).


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