The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 2004. database, Amphibian Species of the World: an Online Reference
  • Duellman, W.E., Lynch, J.D. (1969). ''Description of Atelopus Tadpoles and Their Relevance to the Atelopoid Classication.'' Herpetologica, 25(4), 231-240.
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Lotters, S. (2003). ''On the Systematics of the Harlequin Frogs (Amphibia: Bufonidae: Atelopus) From Amazonia. III: A New, Remarkably Dimorphic Species From the Cordillera Azul, Peru.'' Salamandra, 39(3/4), 169-180.
  • Lötters, Stefan, Salas, Antonio, Angulo, Ariadne, Icochea, Javier, Reynolds, Robert, and La Marca Enrique 2004. Atelopus andinus. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.2. Downloaded on 11 April 2014.
  • Orrell T. (custodian) (2015). ITIS Global: The Integrated Taxonomic Information System (version Mar 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Rivero, J.A. (1968). ''More on the Atelopus (Amphibia, Salientia) from western South America.'' Caribbean Journal of Science, 8(1-2), 19-29.


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