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 Thallus: crustose, of deeply cracked areolate to subsquamulose; areoles: strongly convex, 0.3-0.7 mm thick, from contiguous to dispersed, similar whether sterile or fertile; surface: dark brown or blackish brown; smooth; prothallus: absent; Perithecia: within larger areoles, only mouth showing with part of involucrellum which is thickened only near mouth (subcompound), black or dark brown, merging with thallus tissue; exciple: hyaline to pale brown, blackening toward mouth; asci: saccate, 2-spored; ascospores: ellipsoid or ovoid, dark brown, muriform with many cells, 20-50 x 15-25 µm; Spot tests: all negative; Secondary metabolites: none detected.; Substrate and ecology: on acid or basic rocks often near but not in water; World distribution: circumpolar temperate and boreal; western montane specimens range up to 3600 m and extending to central Mexico; Sonoran distribution: common in Arizona and scattered in southern California and Sonora where limestone occurs and on gneiss, schist and sandstone. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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