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 Thallus: squamulose to small foliose, 1-3 cm diam., often broken up due to growing among mosses, rosette-forming, lobate; lobes: initially rounded, becoming ± flattened and elongate (0.5-1.5 mm wide and up to 5 [-10] mm long), irregularly branched, becoming imbricate; apices: sometimes ascending; upper surface: red-brown when dry, pale brown to deep green when wet, smooth, nodulose or sometimes with globose to cylindrical isidia marginally; upper cortex: paraplectenchymatous, 25-30 micrometer thick; medulla: whitish, loosely interwoven hyphae around Nostoc colonies; lower surface: brownish white, rhizinate; rhizines: brown, sparse, at base of the lobes; Apothecia: not known in Sonoran material, elsewhere rather rare, laminal or marginal, up to 2 mm diam., sessile to substipitate; margin: paler, with short hairs; disc: flat, brown to red-brown, smooth; exciple: 100-145 micrometer wide, true portion paraplectenchymatous, supported by a pseudoexciple; hymenium: brown above, K-, colorless below, I+ blue, 70-130 micrometer; paraphyses: septate simple, 2-6 micrometer wide, ± swollen at the apices and pigmented; subhymenium: hyaline, 45-80 micrometer thick; asci: cylindrical, 55-80 x 11.5-14.5 micrometer, 8-spored; ascospores: fusiform to narrowly ellipsoid, hyaline, 1 (-2)-septate, 11-27 x 4.5-8.5 micrometer; Pycnidia: brown or black above, pale below, immersed, up to 0.6 mm diam.; conidia: bacilliform or slightly bifusiform, colorless, simple, 4-6 x 1 micrometer; Spot tests: all negative; Secondary metabolites: none detected.; Substrate and ecology: among mosses on a range of different acidic rocks in moist habitats at intermediate elevations; World distribution: temperate regions of North and South America, Europe, eastern Asia and Australasia; Sonoran distribution: rare, widely scattered in eastern Arizona. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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