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BiologyOccurs in mountain streams, rivers and lakes (Ref. 41236). Inhabits sandy and gravel-bottomed rivers (Ref. 12045). Herbivore, feeding on bottom detritus (Ref. 40974). Mature adults undertake spawning migration to incoming streams where they breed amidst gravel and sandy beds (Ref. 40974). Fry always occur in quiet parts of the streams or in the side branches of the main streams.