LOBES: distinctly lobate, lobes plane to concave, grayish to pale yellowish-green to pale greenish-yellow, shiny-waxy texture, epruinose or not
 ANATOMY: medulla loose and cottony, algal layer discontinuous
 APOTHECIA: packed in center, discs pale yellowish to orangish to brownish, epruinoes; rim distinct, continuous, like thallus
 CHEMISTRY: cortex usually KC+ gold (usnic acid)