Smilosuchus gregorii was a top predator in a rich environment, as preserved by the Chinle Formation. The Chinle formation is the result of Late Triassic (221.5 to 205.6 million years ago) deposition of aquatic sediments. This could have included fluvial sediments, deposited by by river or stream, lacustrine sediments, deposited by lake, and flood sediments (1). This diverse assemblage contains a range of taxa, including early archosaurs, dinosaurs, phytosaurs, and crocodylomorphs (2). During this time period, the first representatives of many modern groups first appeared in the fossil record, such as dinosaurs and mammals (3). The climate throughout most of the Triassic was very hot and dry, though by the time S. gregorii evolved, there was a shift toward higher levels of humidity (4).
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