Global Range: (20,000-200,000 square km (about 8000-80,000 square miles)) This northeastern North American species was historically found from Nova Scotia to South Carolina in the Atlantic drainages, with an isolated record in Greenbrier River of West Virginia, part of Ohio drainage (Clarke, 1981). Present distribution is spotty, including the Potomac drainage in Virginia, small populations in North and South Carolina, several populations farther north in New York (Neversink River) and elsewhere, and numerous large populations in Maine (Nedeau et al., 2000) and small populations in Massachusetts (Smith, 2000), Connecticut (Nedeau and Victoria, 2003), and New Hampshire (Fichtel and Smith, 1995). Although it was reported in Rhode Island over 100 years ago, there has never been a documented occurrence in the state since (Raithel and Hartenstein, 2006). A discontinuity also exists in northern New Hampshire and southern Maine (COSEWIC, 2009). A relatively large population of a few thousand individuals were recently found in Suncook, New Hampshire (Conaboy, 2006). In Canada, it has a limited distribution in the Bay of Fundy drainage in New Brunswick (8 drainages) and is rare in Nova Scotia (6 drainages) (COSEWIC, 2009).