Adults inhabit lakes and rivers. Juveniles move into the sea and return to rivers after several months. This species has several types differentiated according to shape, coloration, and life history. Each type is isolated by habitat in the same stream.
- Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino 1984 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)
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