The phylogenetics of D. scovazzi are clearly related to the vicariance created by the Strait of Gibraltar and the geological formation of that geographic feature. Morphology and DNA of D. scovazzi have been compared with D. jeanneae. Individuals of the D. scovazzi populations of North Africa and southwest Iberia were sampled and contrasted with sampling of D. jeanneae from eastern Iberia. Morphologies of these two related taxa were differentiated using Mahalanobis distance metrics. Results showed that the effective Mahalanobis distance was greater for the intercontinental than the intracontinental populations by a factor of 2.3 to 3.3. Thus the geographic barrier of the Strait of Gibraltar has had a strong effect on the morphological differentiation as well as the genetic differentiation of the two taxa. Furthermore, mitochondrial DNA and allozyme differentiation is substantial, suggesting the species separation between D. scovazzi and D. jeanneae precedes or is coincident with the formation of the Strait of Gibraltar (Busack & Lawson, 2008).
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