Pseudoceanic, epibenthic species (Ref. 4479, 4775, 51024). Bentho- and mesopelagic in nearshore areas at 0-550 m at night, 500-686 m during day (Ref. 58302).
- Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino 1984 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)
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