The adults spawn in the lake proper at a depth range of 0.5 to 3 m, over fine gravel. Two spawning stocks are known: one spawns in the northwestern corner of the lake from the beginning of November till the end of December; the other spawns at the southeastern corner, from the middle or the end of January till the end of March. This fish approaches the coast twice a year: from April till July and from October till December. In mid winter and at the end of summer, it leaves the coasts to stay at greater depth. Adults feed exclusively on sand hoppers (Amphipoda) (Ref. 593).
- Berg, L.S. 1962 Freshwater fishes of the U.S.S.R. and adjacent countries. volume 1, 4th edition. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1948). (Ref. 593) http://www.fishbase.org/references/FBRefSummary.php?id=593&speccode=4604
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