Inhabits deep parts of large rivers, with moderate to swift current (Ref. 59043). Adults live essentially in freshwater although some fish frequently occur in estuaries. Males are sexually mature between 9-15 years; females between 16-20 years (Ref. 11941). Spawns in main river channel over stone-gravel or gravel-sand bottom and with strong current (Ref. 59043).
- Baillie, J. and B. Groombridge (eds.) 1996 1996 IUCN red list of threatened animals. IUCN, Gland, Switzerland. 378 p. (Ref. 12255) http://www.fishbase.org/references/FBRefSummary.php?id=12255&speccode=4683
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