Oviparous, deposits eggs in small, oval eggs cases on the bottom (Ref. 247). Paired eggs are laid. Embryos feed solely on yolk (Ref. 50449). Distinct pairing with embrace (Ref. 205). During copulation observed in captivity, the male bites the female's pectoral fin in a side-to-side position (Ref. 49562, 51121).
- Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205) http://www.fishbase.org/references/FBRefSummary.php?id=205&speccode=1256
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