Inhabits rubble and gravel riffles of cold creeks and small to medium rivers. Also found in rocky shores of lakes.
- Page, L.M. and B.M. Burr 1991 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723) http://www.fishbase.org/references/FBRefSummary.php?id=5723&speccode=2590
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