Male selects a nesting site under a rock of ledge. Male entices a female to enter the nest. Female deposits eggs on the upper surface of the rock and leaves or is driven out of the nest. Eggs are deposited by different females. The male guards the eggs from predators and maintains a natural current flow.
- Scott, W.B. and E.J. Crossman 1973 Freshwater fishes of Canada. Bull. Fish. Res. Board Can. 184:1-966. (Ref. 1998) http://www.fishbase.org/references/FBRefSummary.php?id=1998&speccode=2594