Demersal spawners. Spawn at night on sand beaches at high tides in the spring and summer. Eggs are buried in moist sand by the female parent (Ref. 58332) and hatch about 15 days later during the next series of high tides (Ref. 2850). Females spawn 4 to 8 times during a season (Ref. 4563).
- Balon, E.K. 1990 Epigenesis of an epigeneticist: the development of some alternative concepts on the early ontogeny and evolution of fishes. Guelph Ichthyol. Rev. 1:1-48.
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