Spawns only during daytime (Ref. 27547). At spawning, males lie close to the bottom in the current of the spawning area while females stay along the banks and in still water (Ref. 27547). A female moves from the bank of the stream usually escorted by 2-4 males to the spawning area at the center of the stream. The males crowd beside her; egg deposition occurs as the males try to clasp the female with their pelvic fins or vibrate against her with their anal fins. This spawning act lasts for 3-5 seconds and may occur as often as 6-40 times per hour. After the eggs are deposited, the sexes separate and return to their previous stream positions (Ref.1998). Fish that moved out of a lake to spawn generally return to the lake a few days after spawning. However, river-resident fish may stay on or near the spawning area for much of the summer (Ref. 27547). Many spawn in two or even three consecutive years but others may skip one or two years between spawning (Ref. 10928).
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