Neustonic to mesopelagic; at the surface at night and to 825 m during day (Ref. 58302).
- Nakabo, T. 2002 Fishes of Japan with pictorial keys to the species, English edition I. Tokai University Press, Japan, pp v-866. (Ref. 41299) http://www.fishbase.org/references/FBRefSummary.php?id=41299&speccode=4669
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