Adults inhabit springs. Mature individuals spawn in February - October (Ref. 59043). Males build, guard and aerate the nest where the eggs are deposited (Ref. 205).
- Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol, Switzerland. 646 p. (Ref. 59043) http://www.fishbase.org/references/FBRefSummary.php?id=59043&speccode=231
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