Inhabits lakes and enters outlet. Spawns at and in outlet of lake in October - January. Potentially extinct due to its original spawning sites are no longer accessible as the outlet of the lake has been obstructed by a weir and two reservoirs have flooded the spawning sites (Ref. 59043).
- Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol, Switzerland. 646 p. (Ref. 59043) http://www.fishbase.org/references/FBRefSummary.php?id=59043&speccode=231
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