Occur mainly in freshwater in large rivers, reservoirs, lakes, swamps, temporary floodwater pools, etc., but adults also found in brackish or saline water of estuaries or bays, preferring quieter open waters. Juveniles are found in great abundance well upstream from brackish water (Ref. 39041). Very young individuals apparently never enter brackish water (Ref. 38947). Larvae are most abundant in surface waters both day and night (Ref. 4639). A herbivorous filter-feeder almost entirely. Breed near the surface in freshwater from late winter (mid-March) through most of the summer (at least to about mid-August). The adhesive eggs sink. Used to some extent as fertilizer and cattle food (Ref. 188).
Whitehead, P.J.P. 1985 FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeioidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/1):1-303. Rome: FAO. (Ref. 188)