Lower jaw slightly projecting, gill cover without any bony radiating striae, teeth rarely present on vomer; belly with a strong keel of scutes; last two anal fin rays not enlarged. No dark spots on flanks. Pterotic bulla absent.
- Whitehead, P.J.P. 1985 FAO Species Catalogue. Vol. 7. Clupeoid fishes of the world (suborder Clupeioidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. FAO Fish. Synop. 125(7/1):1-303. Rome: FAO. (Ref. 188) http://www.fishbase.org/references/FBRefSummary.php?id=188&speccode=24
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